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Finding Aids to Personal Papers and Special Collections in the Smithsonian Institution Archives

Accession 17-132

Heyer, W. Ronald

W. Ronald Heyer Papers, 1963-2015

Repository: Smithsonian Institution Archives, Washington, D.C. Contact us at osiaref@si.edu.
Creator: Heyer, W. Ronald
Title: W. Ronald Heyer Papers
Dates: 1963-2015
Quantity: 15 cu. ft. (15 record storage boxes)
Collection: Accession 17-132
Language of Materials: English
Summary:

This accession consists of papers documenting the career of W. Ronald Heyer, Research Zoologist in the Division of Amphibians and Reptiles at the National Museum of Natural History (NMNH). Heyer's research focuses on the systematics, evolution, and biogeography of Neotropical amphibians. Materials include correspondence, presentations, grant proposals, data, publications, books, maps, posters, scientific illustrations, plates, photographs, negatives, slides, transparencies, and audiovisual materials. Some materials are in electronic format. Some materials were created prior to his tenure at NMNH.

Descriptive Entry

This accession consists of papers documenting the career of W. Ronald Heyer, Research Zoologist in the Division of Amphibians and Reptiles at the National Museum of Natural History (NMNH). Heyer's research focuses on the systematics, evolution, and biogeography of Neotropical amphibians. Materials include correspondence, presentations, grant proposals, data, publications, books, maps, posters, scientific illustrations, plates, photographs, negatives, slides, transparencies, and audiovisual materials. Some materials are in electronic format. Some materials were created prior to his tenure at NMNH.

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Preferred Citation

Smithsonian Institution Archives, Accession 17-132, W. Ronald Heyer Papers

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Container List

Box 1

Correspondence

Adams, Dean C., 2004-2005

Box 1 of 15

Ahlander, Erik, 2003, 2005-2006, 2010-2011, 2015

Box 1 of 15

Allison, Allen, 2006

Box 1 of 15

Almeida, Antonio de Padua, 2006, 2010

Box 1 of 15

Almendariz, Ana, 2012

Box 1 of 15

Altig, Ronald G., 2012

Box 1 of 15

Amaro-Ghilardi, Renata Cecilia, 2002

Box 1 of 15

Ananjeva, Natalia, 2002

Box 1 of 15

Anderson, Karen, 1981, 1995

Box 1 of 15

Annunziata, Bruno Barcellos, 2007

Box 1 of 15

Anstis, Marion, 2005-2006, 2008

Box 1 of 15

Araujo, Olivia Gabriela, 2009, 2013

Box 1 of 15

Arntzen, J. W., 2004, 2007

Box 1 of 15

Arruda, Mauricio Papa de, 2006

Box 1 of 15

Arzabe, Cristina, 2003

Box 1 of 15

Avila Pires, Teresa, 2004, 2009

Box 1 of 15

Ayarzaguena, Jose, 1990

Box 1 of 15

Badillo, Alberto Fernandez, 1990

Box 1 of 15

Baeta, Delio, 2010

Box 1 of 15

Ballentine, Barbara, 2009

Box 1 of 15

Barg, Monica, 2001

Box 1 of 15

Barrio Amoros, Cesar Luis, 2011-2013

Box 1 of 15

Basso, Nestor, 2009

Box 1 of 15

Bauer, Aaron M., 2008-2009

Box 1 of 15

Berneck, Bianca, 2008

Box 1 of 15

Biancardi, Carlo, 2009

Box 1 of 15

Bickford, David, 2010

Box 1 of 15

Bilate, Marcos, 2008

Box 1 of 15

Bionda, Clarisa de Lourdes, 2009

Box 1 of 15

Bisbal, Francisco, 1990

Box 1 of 15

Bogart, Jim, 2006

Box 1 of 15

Bohme, Wolfgang, 2003

Box 1 of 15

Boistel, Renaud, 2001, 2005

Box 1 of 15

Briggs, Jeffrey, 2005

Box 1 of 15

Bronikowski, Ed, 2008

Box 1 of 15

Bryan, Ann Wascher, 2011

Box 1 of 15

Burge, Russell, 2004

Box 1 of 15

Bustamante, Martin R., 2008

Box 1 of 15

Caldwell, Janalee P., 2004, 2006, 2008-2010

Box 1 of 15

Cannatella, David, 2002-2003, 2006-2007

Box 1 of 15

Caramaschi, Ulisses, 1997, 2000-2004, 2006-2009

Box 1 of 15

Cardoso, Manoela, 2010

Box 1 of 15

Cardozo, Martin, 2004-2005

Box 1 of 15

Carstensen, Daniel Wisbech, 2006

Box 1 of 15

Carvalho, Celso Morato de, 1998, 2008-2012

Box 1 of 15

Carvalho, Thiago Ribeiro de, 2012, 2015

Box 1 of 15

Cascon, Paulo, 2008

Box 1 of 15

Cei, Jose M., 2004

Box 1 of 15

Chestnut, Tara, 2009

Box 1 of 15

Cisneros-Heredia, Diego F., 2006

Box 1 of 15

Cocroft, Rex, 2006

Box 1 of 15

Coddington, Jonathan, 2012

Box 1 of 15

Cole, Charles J., 2003, 2007-2010

Box 1 of 15

Colli, Guarino R., 2003, 2009-2010

Box 1 of 15

Cordova Santa Gadea, Jesus H., 2006

Box 1 of 15

Costa, Henrique Caldeira, 2009, 2012

Box 1 of 15

Cox, Neil, 2006-2007

Box 1 of 15

Crawford, Andrew J., 2006, 2010

Box 1 of 15

Crump, Martha, 2005

Box 1 of 15

Cruz, Carlos Alberto, 2012-2013

Box 1 of 15

Curcio, Felipe Franco, 2009

Box 1 of 15

Czapracki, Allison, 2009

Box 1 of 15

David, Patrick, 2004

Box 1 of 15

de Queiroz, Kevin, 2007

Box 1 of 15

de Ruiter, Ronald, 2009

Box 1 of 15

de Sa, Rafael, 2008-2015

Box 1 of 15

Dias, Iuri Ribeiro, 2008, 2011

Box 1 of 15

Dixon, James R., 1998, 2000, 2002, 2005, 2007

Box 1 of 15

Dodd, Kenneth, 2009

Box 1 of 15

Donnelly, Maureen A., 2009

Box 1 of 15

Drewery, George

Box 1 of 15

Duarte B. Filho, Jose, 2009

Box 1 of 15

Dundee, Harold, 2009

Box 1 of 15

Erdtmann, Luciana, 2008

Box 1 of 15

Ernst, Carl H., 2013

Box 1 of 15

Espanola, Asociacion Herpetologica, 2008

Box 1 of 15

Eterovick, Paula, 2004-2005, 2007

Box 1 of 15

Fabrezi, Marissa, 2007

Box 1 of 15

Faivovich, Julian, 2002, 2010

Box 1 of 15

Feio, Renato, 2008

Box 1 of 15

Felsenstein, Joseph, 1976

Box 1 of 15

Fong G., Ansel, 2007

Box 1 of 15

Ford, Linda S., 1979-1980, 1988, 1990, 1992, 1998

Box 1 of 15

Forlani, Mauricio da Cruz, 2009, 2012-2013

Box 1 of 15

Fouquet, Antoine, 2009-2010, 2012

Box 1 of 15

Frost, Darrel, 1997, 2005-2006, 2008, 2012

Box 1 of 15

Franklin, Carl J., 2010

Box 1 of 15

Freitas, Joseana, 2011

Box 1 of 15

Gagliardi Urrutia, Giuseppe, 2010

Box 1 of 15

Garcia, Paulo C. A., 2011, 2013

Box 1 of 15

Garda, Adrian Antonio, 2009

Box 1 of 15

Gascon, Claude, 2011

Box 1 of 15

Ghilardi, Renata Cecilia Amaro, 2004

Box 1 of 15

Giaretta, Ariovaldo Antonio, 2006-2008, 2011

Box 1 of 15

Giasson, Luis Menta, 2007

Box 1 of 15

Gibula, Jeff, 2008

Box 1 of 15

Good, Tatjana, 2007

Box 1 of 15

Gordo, Marcelo, 2007

Box 1 of 15

Gorzula, Stefan, 1990

Box 1 of 15

Grant, Taran, 2006-2007, 2009, 2013

Box 1 of 15

Green, David E., 2001

Box 1 of 15

Guayasamin, Juan Manuel, 2005

Box 1 of 15

Guerreiro Cordova, Manuel Alejandro, 2008

Box 1 of 15

Haddad, Celio F. B., 1993, 2003-2004

Box 1 of 15

Hanken, James, 2006

Box 1 of 15

Hansen, Robert W., 2013

Box 1 of 15

Hartsell, Traci D., 2009

Box 1 of 15

Hawley, Tanya, 2007

Box 1 of 15

Hayek, Lee-Ann, 2001

Box 1 of 15

Henderson, Robert, 2006

Box 1 of 15

Henle, Klaus, 2003

Box 1 of 15

Hepp, Fabio Silva Fernandes dos Santos, 2012

Box 1 of 15

Hero, Jean-Marc, 2007, 2010

Box 1 of 15

Herrera Villalobos, Alvaro, 2006-2007

Box 1 of 15

Hibbitts, Toby, 2005, 2007, 2010

Box 1 of 15

Hicks, Tracy

Box 1 of 15

Hoogmoed, Marinus, 2011-2012, 2014

Box 1 of 15

Hoyer, Richard F., 2004

Box 1 of 15

Hurme, Kristiina J., 2004, 2012-2014

Box 1 of 15

Ibanez, Roberto, 2005, 2007

Box 1 of 15

Inga, Jorge Joel, 2009

Box 1 of 15

Inger, Robert F., 2010

Box 1 of 15

Jackson, Kate, 2009

Box 1 of 15

Jayne, Bruce C., 1988

Box 1 of 15

Johnson, G. David, 2007

Box 1 of 15

Johnson, Susan, 2003

Box 1 of 15

Joventino Roberto, Igor, 2006, 2014

Box 1 of 15

Jowers, Michael, 2003

Box 1 of 15

Junca, Flora Acuna, 2000-2002, 2004-2005, 2011

Box 1 of 15

Jungfer, Karl-Heinz, 2009

Box 1 of 15

Kaiser, Hinrich, 2005

Box 1 of 15

Kaplan, Moises, 2004

Box 1 of 15

Kattan, Gustavo, 2008

Box 1 of 15

King, Jay, 2012

Box 1 of 15

Kizirian, David, 2007

Box 1 of 15

Klaczko, Julia, 2005

Box 1 of 15

Koehler, Gunther, 2011-2012

Box 1 of 15

Kok, Philippe, 2006

Box 1 of 15

Kokubum, Marcelo N. de C., 2013

Box 1 of 15

Kozak, Kenneth H., 2007

Box 1 of 15

Kretzschmar, Sonia, 2003, 2010

Box 1 of 15

Kullander, Sven O., 2003

Box 1 of 15

Kwet, Axel, 2004-2005, 2007

Box 1 of 15

Lacerda, Joao Victor A. de, 2009

Box 1 of 15

La Marca, Enrique, 2011

Box 1 of 15

Langstroth, Robert, 2010

Box 1 of 15

Lannoo, Michael J., 2002

Box 1 of 15

Lazell, Skip, 2005

Box 1 of 15

Lee, David, 2008

Box 1 of 15

Leite, Felipe S. F., 2008

Box 1 of 15

Lips, Karen, 2008-2009

Box 1 of 15

Lobo, Fernando, 2007

Box 1 of 15

Loebmann, Daniel, 2006

Box 1 of 15

Loman, Jon, 2009

Box 1 of 15

Losos, Jonathan B., 2000

Box 1 of 15

Lutz, Adolfo

Box 1 of 15

Luhring, Thomas M., 2007

Box 1 of 15

Lynch, John D., 2004, 2013, 2015

Box 1 of 15

MacCulloch, Ross, 2006

Box 1 of 15

Magnusson, William, 2008

Box 1 of 15

Manzanilla, Jesus, 2006

Box 1 of 15

Marquez, Rafael, 2011

Box 1 of 15

Marx, Hymen, 2007

Box 1 of 15

Maxson, Linda E. R., 1979-1980, 2007

Box 1 of 15

May, Heather A., 2008

Box 1 of 15

Mayer, Gregory C., 2006

Box 1 of 15

McCracken, Shawn F., 2009

Box 1 of 15

McCranie, James R., 2012

Box 1 of 15

McDiarmid, Roy W., 2008

Box 1 of 15

McGuire, Jimmy A., 2007

Box 1 of 15

McLeod, David S., 2005

Box 1 of 15

McQuoid-Greason, Meredith, 2011

Box 1 of 15

Medina, Regina Gabriela, 2013

Box 1 of 15

Mello, Caio Marinho, 2014

Box 1 of 15

Mello, Carolina, 2009, 2011

Box 1 of 15

Melo, Paulo, 2011

Box 1 of 15

Mendelson, Joseph R., III, 2008

Box 1 of 15

Mendoza Roldan, Juan Salvador, 2014

Box 1 of 15

Meneghelli, Diego, 2011-2012

Box 1 of 15

Mijarres, Abraham, 1999, 2011

Box 1 of 15

Minter, Leslie, 2006

Box 1 of 15

Mitchell, Joseph C., 2007

Box 1 of 15

Moen, Daniel S., 2011

Box 1 of 15

Moler, Paul, 2002

Box 1 of 15

Molur, Sanjay, 1999

Box 1 of 15

Morgan, Gary S., 2009

Box 1 of 15

Moro, Silvia, 1996

Box 1 of 15

Munoz, Arturo, 2005

Box 1 of 15

Murphy, John C., 2011

Box 1 of 15

Myers, Charles W., 2011

Box 1 of 15

Nascimento, Luciana Barreto, 2005, 2007, 2010

Box 1 of 15

Nomura, Hitoshi, 2004

Box 1 of 15

Nunes, Ivan, 2006, 2008, 2013, 2015

Box 1 of 15

Ohler, Annemarie, 2009

Box 1 of 15

Oliveira, Leonardo de, 2010

Box 1 of 15

Orrico, Victor G. Dill, 2010

Box 1 of 15

Ortega-Andrade, H. Mauricio, 2010

Box 1 of 15

Ospina-Sarria, Jhon Jairo, 2012

Box 1 of 15

Ouboter, Paul E., 2010

Box 1 of 15

Oyamaguchi, Hilton, 2008

Box 1 of 15

Padial, Jose, 2011

Box 1 of 15

Paez V., Monica, 2006

Box 1 of 15

Parham, James, 2007

Box 1 of 15

Pase, H. A. (Joe), III, 2010

Box 1 of 15

Passos, Paulo, 2004

Box 1 of 15

Pauly, Gregory B., 2004, 2014

Box 1 of 15

Pauwels, Olivier S. G., 2008

Box 1 of 15

Peao, Carlos, 2007

Box 1 of 15

Pearson, David L., 2003

Box 1 of 15

Peloso, Pedro L. V., 2011

Box 1 of 15

Box 2

Peltzer, Paola, 2004

Box 2 of 15

Peres, Juliana, 2008

Box 2 of 15

Perry, Gad, 2006

Box 2 of 15

Peters, Gustav, 2005

Box 2 of 15

Pinedo, Jorge Joel, 2010

Box 2 of 15

Pinto, Gabriel Silva, 2000, 2004

Box 2 of 15

Pombal, Jose P., Jr., 2011

Box 2 of 15

Ponssa, Maria Laura, 2005, 2011

Box 2 of 15

Pontes, Rafael Cunha, 2010-2011

Box 2 of 15

Prado, Cynthia P. A., 2005

Box 2 of 15

Prado, Gustavo M., 2005

Box 2 of 15

Pregill, Gregory K., 2007

Box 2 of 15

Provete, Diogo Borges, 2008

Box 2 of 15

Prudente, Ana, 2007-2008

Box 2 of 15

Rapp, Tain , 2008

Box 2 of 15

Relyea, Rick, 2009, 2011

Box 2 of 15

Resetar, Alan, 2008

Box 2 of 15

Ribeiro, Joana, 2014

Box 2 of 15

Ribeiro, Marco Antonio, Jr., 2010

Box 2 of 15

Rice, Jeff, 2007

Box 2 of 15

Roberto, Igor Joventino, 2005-2006

Box 2 of 15

Robertson, Jeanne M., 2005-2006

Box 2 of 15

Robinson, Earle, 2006

Box 2 of 15

Rocek, Zbynek, 2005

Box 2 of 15

Rocha, Carlos Frederico Duarte da, 2006

Box 2 of 15

Rocha, Mariane Targino, 2006, 2008-2009

Box 2 of 15

Rodriguez Bayona, Lily Olga, 2010-2011

Box 2 of 15

Roedel, Mark-Oliver, 2007

Box 2 of 15

Roelofs, Terry, 2010

Box 2 of 15

Romero, Heron, 2006

Box 2 of 15

Ron, Santiago, 2005-2006, 2011

Box 2 of 15

Rosado, Jose, 2011

Box 2 of 15

Rossman, Doug, 2008

Box 2 of 15

Samper, Cristian, 2007, 2009

Box 2 of 15

Samyn, Yves, 2008-2009

Box 2 of 15

Sanchez Ramirez, David, 2004-2005

Box 2 of 15

Sanchez Pacheco, Santiago J., 2009

Box 2 of 15

Santos, Jose W. A., 2008

Box 2 of 15

Santos, Juan C., 2006

Box 2 of 15

Santos, Marcus Thadeu Teixeira, 2013

Box 2 of 15

Savage, Jay M., 2012

Box 2 of 15

Savitzky, Alan H., 2008-2009

Box 2 of 15

Sazima, Ivan, 1980, 2003, 2007, 2014

Box 2 of 15

Scharlach, Cecilia, 2009

Box 2 of 15

Schiesari, Luis, 2005, 2008

Box 2 of 15

Schmid, Michael, 2005

Box 2 of 15

Schulze, Arne, 2010

Box 2 of 15

Segalla, Magno, 2009, 2011

Box 2 of 15

Senaris, J. Celsa, 2011

Box 2 of 15

Seripierri, Dione, 2002, 2006

Box 2 of 15

Sheridan, Jennifer, 2005-2006

Box 2 of 15

Silva, Helio Ricardo da, 2003, 2006, 2011

Box 2 of 15

Silva-Filho, Ivan Sergio N., 2006

Box 2 of 15

Silva, Monica Carolina Cardoso da, 2008-2009

Box 2 of 15

Silva-Soares, Thiago, 2012-2013

Box 2 of 15

Simmons, John E., 2004

Box 2 of 15

Simon, Monique N., 2012-2013

Box 2 of 15

Slough, Brian G., 2003

Box 2 of 15

Smith, Joanna, 2011

Box 2 of 15

Smith, Paul, 2011

Box 2 of 15

Smythe, Ashleigh, 2008

Box 2 of 15

Solano de Chacin, Haydee, 1990

Box 2 of 15

Sole, Mirco, 2006-2007, 2010-2011

Box 2 of 15

Souza, Sergio Marques de, 2008

Box 2 of 15

Spencer, Kate, 1994

Box 2 of 15

Stafford, Peter J., 2007

Box 2 of 15

Starr, Christopher K., 2005

Box 2 of 15

Strader, Ginger, 2009

Box 2 of 15

Strussmann, Christine, 2004

Box 2 of 15

Stuart, Simon N., 2006, 2008

Box 2 of 15

Tarano, Zaida, 2011

Box 2 of 15

Teixeira Junior, Mauro, 2011

Box 2 of 15

Teixeira, Rogerio, 2005

Box 2 of 15

Tol, J. van, 2007

Box 2 of 15

Torres, Omar, 2008

Box 2 of 15

Torres-Cervantes, Ricardo, 2005

Box 2 of 15

Trueb, Linda, 2006, 2008

Box 2 of 15

Uzzell, Thomas, 2009

Box 2 of 15

Val, Aldoberto, 2006

Box 2 of 15

Val, Francisca Carolina (Chica) do, 1991, 1994, 1997-2015

Box 2 of 15

Vanzolini, Paulo Emilio, 2013

Box 2 of 15

Vari, Richard, 2012

Box 2 of 15

Veauvy, Mariana, 2008

Box 2 of 15

Velez R., Caudia Maria, 2004

Box 2 of 15

Verdade, Vanessa K., 2004-2007

Box 2 of 15

Vieira, Kleber S., 2007, 2009

Box 2 of 15

Vigle, Greg, 2006

Box 2 of 15

Vogt, Richard, 2005-2008

Box 2 of 15

Voris, Harold K., 2008

Box 2 of 15

Wake, David, 2007-2008

Box 2 of 15

Wake, Marvalee H., 2005, 2007

Box 2 of 15

Waldez, Fabiano, 2007

Box 2 of 15

Walker F. de Faria, Marina, 2012

Box 2 of 15

Walley, Harlan D., 2007

Box 2 of 15

Wassersug, Richard, 2006, 2009

Box 2 of 15

Weigt, Lee, 2010

Box 2 of 15

Williams, Jorge, 2005, 2009

Box 2 of 15

Wilson, Don, 2009

Box 2 of 15

Wray, Ken, 2008

Box 2 of 15

Wynn, Addison, 2011, 2014

Box 2 of 15

Yanez-Munoz, Mario Humberto, 2006, 2009

Box 2 of 15

Zaher, Hussam, 2004-2006, 2008-2013

Box 2 of 15

Zamudio, Kelly R., 2004-2006, 2012

Box 2 of 15

Zimmerman, Barbara Lewis, 1991, 2001

Box 2 of 15

Research and Manuscripts

Data for presentation at Annual Meeting of the Society of Integrative and Comparative Biology, Patras, Greece, 2002

Box 2 of 15

Leptodactylus troglodytes call analysis for 1995 manuscript

Box 2 of 15

Presentation: 246 years of Nomenclatural Novelties in the Genus Leptodactylus: Past, Present, Future

Box 2 of 15

Brazilian tadpoles, data and illustrations

Box 2 of 15

Brazilian tadpoles, references

Box 2 of 15

Leptodactylus gazetteer

Box 2 of 15

Smithsonian Amphibian Initiative, 2008

Box 2 of 15

Texas collections trip, March 2010

Box 2 of 15

National Public Radio "Talk of the Nation: Science Friday" presentation on declining amphibian problem, 5/16/1997 (audiocassette)

Box 2 of 15

Box 3

Leptodactylus validus manuscript and data, Proceedings of the Biological Society of Washington, 125(2): 276-294, 2012

Box 3 of 15

Leptodactylus annotated bibliography, circa 1965- : A-B

Box 3 of 15

Leptodactylus annotated bibliography, circa 1965- : C-E

Box 3 of 15

Leptodactylus annotated bibliography, circa 1965- : F-H

Box 3 of 15

Leptodactylus annotated bibliography, circa 1965- : I-lutzi

Box 3 of 15

Leptodactylus annotated bibliography, circa 1965- : macroblepharus-notoaktites

Box 3 of 15

Leptodactylus annotated bibliography, circa 1965- : occidentalis-pentadactylus

Box 3 of 15

Leptodactylus annotated bibliography, circa 1965- : perlaevis-R

Box 3 of 15

Leptodactylus annotated bibliography, circa 1965- : S-wuchereri

Box 3 of 15

L. fuscus group standards

Box 3 of 15

L. latrans group standards

Box 3 of 15

L. pentadactylus group standards

Box 3 of 15

Variation, Systematics, and Relationships of the Leptodactylus bolivianus Complex, 2011 (final draft)

Box 3 of 15

Paulo Emilio Vanzolini background obituary materials and drafts, 2013

Box 3 of 15

L. pentadactylus biogeography project: Statistics and data

Box 3 of 15

L. insularum manuscript, Proceedings of the Biological Society of Washington, 126 (3): 204-233, 2013 (2 folders)

Box 3 of 15

Box 4

L. insularum bibliography, Proceedings of the Biological Society of Washington, 126 (3): 204-233, 2013

Box 4 of 15

FAPESP (Sao Paulo Research Foundation) proposal, Leptodactylus at the Museu de Zoologia da Universidade de Sao Paulo (MZUSP), 2010

Box 4 of 15

Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, northern South America, and Amazonia, Arquivos de Zoologia, 37(3): 269-348, 2005 - Measurement data coding

Box 4 of 15

Leptodactylus pentadactylus, Catalogue of American Amphibians and Reptiles, (888): 1-47, 2011 - Correspondence, figures, etc.

Box 4 of 15

Leptodactylus pentadactylus, Catalogue of American Amphibians and Reptiles, (888): 1-47, 2011 - Data, drafts, annotated references (5 folders)

Box 4 of 15

L. latrans group data sheets

Box 4 of 15

L. latrans complex, photographs/data - Argentina: San Juan, San Luis, Santa Fe, Santiago del Estero

Box 4 of 15

Box 5

L. latrans complex, photographs/data - Argentina: Tucuman

Box 5 of 15

L. latrans complex, data sheets - Argentina: Buenos Aires, Chaco, Cordoba, Corrientes, Entre Rios, Formosa, Jujuy, La Pampa, Mendoza, Misiones, Salta, San Juan, San Luis, Santa Fe, Santiago del Estero, Tucuman

Box 5 of 15

L. latrans complex, photographs/data - Bolivia: Beni, Chuquisaca, La Paz

Box 5 of 15

L. latrans complex, photographs/data - Bolivia: Santa Cruz

Box 5 of 15

L. latrans complex, photographs/data - Bolivia: Beni, Chuquisaca, La Paz, Santa Cruz

Box 5 of 15

L. latrans complex, photographs/data - Brasil: Alagoas, Amapa, Amazonas, Bahia, Ceara, Distrito Federal, Espirito Santo

Box 5 of 15

L. latrans complex, photographs/data - Brasil: Amazonas - Coari

Box 5 of 15

L. latrans complex, photographs/data - Argentina: No locality, Distrito Buenos Aires, Chaco

Box 5 of 15

L. latrans complex, photographs/data - Argentina: Cordoba, Corrientes

Box 5 of 15

Box 6

L. latrans complex, photographs/data - Argentina: Entre Rios, Formosa

Box 6 of 15

L. latrans complex, photographs/data - Argentina: Jujuy, La Pampa, Mendoza, Misiones, Neuquen, Salta

Box 6 of 15

L. latrans complex, photographs/data - Brasil: Amazonas - Coari

Box 6 of 15

L. latrans complex, photographs/data - Brasil: Goias

Box 6 of 15

L. latrans complex, photographs/data - Brasil: Maranhao, Mato Grosso

Box 6 of 15

L. latrans complex, photographs/data - Brasil: Mato Grosso do Sul

Box 6 of 15

L. latrans complex, photographs/data - Brasil: Minas Gerais

Box 6 of 15

L. latrans complex, photographs/data - Brasil: Para, Paraiba, Pernambuco, Piaui, Rio de Janeiro

Box 6 of 15

L. latrans complex, photographs/data - Brasil: Rio Grande do Norte, Rondonia, Roraima, Santa Catarina, Sao Paulo, Sergipe, Tocantins

Box 6 of 15

L. latrans complex, data sheets - Brasil: Acre, Alagoas, Amapa, Amazonas - Coari, Bahia, Ceara, Distrito Federal, Goias

Box 6 of 15

Box 7

L. latrans complex, data sheets - Brasil: Maranhao, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Para, Paraiba, Parana, Pernambuco, Piaui

Box 7 of 15

L. latrans complex, data sheets - Brasil: Rio de Janeiro, Rio Grande do Sul, Rondonia, Roraima, Santa Catarina, Sao Paulo, Sergipe, Tocantins

Box 7 of 15

L. latrans complex, photographs/data - Guyana; Venezuela: Apure, Bolivar

Box 7 of 15

L. latrans complex, photographs/data - Venezuela: Guarico

Box 7 of 15

L. latrans complex, data sheets - Guyana; Venezuela: Apure, Bolivar, Guarico

Box 7 of 15

L. latrans complex, photographs/data - Trinidad

Box 7 of 15

L. latrans complex, photographs/data - Paraguay: Alto Paraguay, Amambay, Boqueron, Canindeyu, Central, Cordillera, Itapua

Box 7 of 15

L. latrans complex, photographs/data - Paraguay: Paraguari, Presidente Hayes, San Pedro

Box 7 of 15

L. latrans complex, photographs/data - Uruguay: Canelones, Maldonado, Rivera, Rocha, Treinta y Tres

Box 7 of 15

L. latrans complex, data sheets - Paraguay, Uruguay

Box 7 of 15

L. latrans complex - Preliminary discriminant function analyses

Box 7 of 15

L. latrans complex - Miscellaneous data

Box 7 of 15

L. latrans complex - Analyses from digital photographs (as L. ocellatus)

Box 7 of 15

L. latrans complex - Regional analyses of morphological data manuscript

Box 7 of 15

L. latrans complex - Discriminant function analyses data, draft, regional samples

Box 7 of 15

L. latrans complex - Manuscript, various documents

Box 7 of 15

Box 8

Slides of Brasilian field work sites and animals

Box 8 of 15

Dissertation plates

Box 8 of 15

Publication illustrations, originals (includes electronic records)

Box 8 of 15

Leptodactylus validus poster presentation

Box 8 of 15

"The Last Frog" National Geographic presentation (includes VHS)

Box 8 of 15

Howard University presentation - "Why Study Frogs from Nature?", 1989

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"Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians," online chapters

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"Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians," 1994 (English)

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"Medicion y Monitoreo de la Diversidad Biologica: Metodos estandarizados para Anfibios," 1994 (Spanish)

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"Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians" (Russian)

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Amazon presentation - Curitiba herpetology meetings

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Heyer and de Sa National Science Foundation (NSF) proposal, 1996-1997

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Heyer and de Sa NSF award 9815787

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Heyer and de Sa NSF proposal, 2003

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Box 9

Heyer and de Sa NSF grant records and finances as of July 2004

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Leptodactylus phylogeny data and correspondence

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Leptodactylus phylogeny monograph final publication, South American Journal of Herpetology Special Issue, 9 (S1-S128), 2014

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Leptodactylus phylogeny monograph drafts and data (6 folders)

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Leptodactylus phylogeny frog call recordings and requests, photographs and requests

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Leptodactylus phylogeny tissue data and POY runs

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Leptodactylus phylogeny tree

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Leptodactylus phylogeny tissues data

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Leptodactylus phylogeny monograph correspondence

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Cornell Laboratory of Ornithology NSF proposal for frog recordings

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Leptodactylus website common names, calls, photographs

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Box 10

Unpublished Leptodactylus sexual dimorphism manuscript, 2015

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Sexual dimorphism data analysis: Leptodactylus albilabris - Leptodactylus discodactylus

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Sexual dimorphism data analysis: Leptodactylus elenae - Leptodactylus insularum

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Sexual dimorphism data analysis: Leptodactylus knudseni - Leptodactylus mystacinus

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Sexual dimorphism data analysis: Leptodactylus natalensis - Leptodactylus pustulatus

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Sexual dimorphism data analysis: Leptodactylus rhodomerus - Leptodactylus turimiquensis

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Sexual dimorphism data analysis: Leptodactylus validus - Leptodactylus wagneri

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Sexual dimorphism manuscript initial drafts (includes electronic records)

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Sexual dimorphism manuscript data and correspondence

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Seminar presentations notes

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Presentation slides: "Relationships of Leptodactylus silvanimbus"

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Presentation slides: Wynn and Heyer, "Leptodactylus fuscus, one or more species"

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Presentation slides: "Speciation factors in Leptodactylus fuscus, Leptodactylus pentadactylus, and Leptodactylus melanonotus groups"

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Heyer manuscripts (electronic records)

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Leptodactylus pdf files (electronic records)

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Estuarium frog talk (electronic records)

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Speciation patterns in L. pentadactylus cluster (electronic records)

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Brazilian Herpetological Society Belo Horizonte presentation (electronic records)

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Leptodactylus bolivianus complex presentation (electronic records)

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Box 11

EndNote presentation, Leptodactylus research topics, Rio de Janeiro and San Juan (electronic records)

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Belem, Brazil, presentation (electronic records)

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Leptodactylus systematics (electronic records)

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Argentina presentation (electronic records)

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7th Congress, Argentina, presentation (electronic records)

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San Luis, Argentina, presentation (electronic records)

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Locality records - Thailand

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Locality data and correspondence - South America

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Leptodactylus locality data (2 folders)

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Leptodactylus locality Gazetteer

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Drawings, sonograms, maps, data, and photographs for publication

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56 Years of Leptodactylus Data

Marmoratus group - locality data

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Venezuelan Leptodactylus species data

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Leptodactylus locality data

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JAP specimens to borrow

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Negatives

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External morphology data

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Leptodactylus - Tadpole data

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Barycholos

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Lithodytes

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External morphology of melanonotus group

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Leptodactylus - Egg and egg deposition information

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Leptodactylus - Taxonomic notes and miscellaneous

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Chromosome analysis

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L. labialis - Analyses

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L. albilabris - Analyses

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Leptodactylus locality data

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Fuscus group data

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Mystaceus - SW complex form

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Mystaceus

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Fuscus group - Nomenclature correspondence

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Completed data sheets

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Pentadactylus group nomenclature

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Box 12

Leptodactylus data sheets

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Leptodactylus - Breeding call information

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Leptodactylid literature

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Chromosome data

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Heyer and Pyburn - Leptodactylus riveroi

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Pattern standards - Melanonotus group

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Post podicipinus-wagneri manuscript locality data

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Leptodactylus - Distributional data post revisions

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Cycloramphus negatives

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Centrolenella negatives

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Negatives for fuscus group paper

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Map negatives for fuscus group paper

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Proof sheet and negatives for figures associated with L. rugosus, etc., manuscript

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Leptodactylus paraguayensis and L. didymus negatives

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Map negative of South America

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Leptodactylus type negatives

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E. guentheri negatives

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Podicipinus-wagneri complex negatives

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Large species of Leptodactylus holotype negatives and proof sheet

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Eleuth. dundeei, Rupirana cardosoi, Rondonia frog negatives

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Unidentified negatives (2 folders)

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Fuscus group standard sheets

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Fuscus group analyses and notes for manuscript

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Tepequem collection notes

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Cadle Bolivian frog

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Types - Senckenberg, Vienna, Munich, September 1982

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Leptodactylus types A-L

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Leptodactylus types M-Z

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L. delattini

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L. cf furnarius - Pico das Almas

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L. ocellatus

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L. species "elenae" from Peru

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Albilabris - Data and transformations

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L. bolivianus

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L. bufonius

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L. caatingae

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L. camaquara

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L. chaquensis

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L. colombiensis

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L. cunicularius

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L. cupreus

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L. didymus - Data

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L. diedrus

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Vanzolinius discodactylus

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L. elenae

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L. fallax

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L. flavopictus

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L. furnarius

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L. fragilis

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Box 13

Fuscus - Data

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L. jolyi

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L. knudseni

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L. labrosus

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L. labyrinthicus

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L. laticeps

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L. latinasus

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L. gracilis

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L. griseigularis

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L. insularum

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L. lauramiriamae

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L. leptodactyloides

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L. lithonaetes

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L. longirostris

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L. macrosternum

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L. magistris

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L. marambaiae

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L. melanonotus

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L. myersi

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L. mystaceus

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L. mystacinus

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L. natalensis

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L. nesiotus

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L. notoakfites

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L. ocellatus

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L. paraensis

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L. pascoensis

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L. pentadactylus - Amazonia

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L. peritoaktites

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L. petersii

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L. plaumanni

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L. podicipinus

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L. poecilochilus

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L. pustulatus

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Ceara L. pustulatus

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L. rhodomerus

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L. rhodomystax

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L. rhodonotus

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L. riveroi

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L. rugosus

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L. sabanensis

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L. savagei

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L. silvanimbus

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L. spixi

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L. stenodema

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L. syphax

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L. tapiti

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L. troglodytes

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L. turimiquensis

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L. validus

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L. vastus

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L. ventrimaculatus

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L. viridis

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L. wagneri

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Physalaemus gracilis

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Completed melanonotus distribution data sheets

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Distribution notes on the species groups

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Distribution of melanonotus group

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Underwater Component in the Call of the Frog Leptodactylus ocellatus, by Alejandro Purgue

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Caldwell - Brasilian frog data - NSF reports

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J. R. Dixon data on Leptodactylus at El Bagual, Argentina

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Chiasmocleis

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Call of Eleutherodactylus sp. Brejo da Lapa, WRH 8429

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Aborted Leptodactylus pentadactylus biogeography project

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Wynn and Heyer - Genetic relatedness within L. fuscus (electronic records)

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Bertha Lutz bibliography

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Brasil frog literature, notes, etc.

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Leptodactylus loan sheets - Completed

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Leptodactylus types - Negatives and miscellaneous

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Current loans

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Leptodactylus labialis - Distribution

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Pentadactylus group - Locality information

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Rana macrocephala Wied (=Ceratrophrys)

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Box 14

The Society of the Sigma Xi certificate, 1966

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Phi Sigma Society certificate, 1964

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"A Diatom Is Two Atoms Working Together" - A collection of classic examination answers of biology 100 and 106 students

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Leptodactylus - Miscellaneous correspondence, 1963-1968

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Completed Pacific Lutheran University loan invoices 1-25

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Leptodactylus correspondence and miscellaneous (3 folders)

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Leptodactylus data for phylogenetic analysis

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Leptodactylus holdings at University of Texas, Arlington, 2006

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Anais da Academia Brasileira de Ciencias, 76(2), 2004

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Publications

Heyer bibliography

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Heyer, W. R. 1967. A herpetofaunal study of an ecological transect through the Cordillera de Tilaran, Costa Rica. Copeia, 1967: 259-271.

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Savage, J. M., and Heyer, W. R. 1967. Variation and distribution in the tree-frog genus Phyllomedusa in Costa Rica, Central America. Beitrage zur Neotropischen Fauna, 5: 111-131.

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Dixon, J. R., and Heyer, W. R. 1968. Anuran succession in a temporary pond in Colima, Mexico. Bulletin of the Southern California Academy of Sciences, 67: 129-137.

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Heyer, W. R. 1968. The proper name for the type-species of the genus Leptodactylus. Copeia, 1968: 160-162.

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Heyer, W. R. 1969. The adaptive ecology of the species groups of the genus Leptodactylus (Amphibia, Leptodactylidae). Evolution, 23: 421-428.

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Heyer, W. R., and Silverstone, P. A. 1969. The larva of the frog Leptodactylus hylaedactylus (Leptodactylidae). Fieldiana Zoology, 51: 141-145.

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Heyer, W. R. 1969. Studies on frogs of the genus Leptodactylus (Amphibia, Leptodactylidae). 5. Taxonomic notes on L. latinasus, rhodonotus, romani and wuchereri. Herpetologica, 25: 1-8.

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Heyer, W. R. 1969. Studies on the genus Lepodactylus (Amphibia, Leptodactylidae). 3. A redefinition of the genus Leptodactylus and a description of a new genus of leptodactylid frogs. Contributions in Science, 155: 1-14.

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Savage, J. M., and Heyer, W. R. 1969. The tree-frogs (Family Hylidae) of Costa Rica: Diagnosis and distribution. Revista de Biologia Tropical, 16(1): 1-127.

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Heyer, W. R. 1970. Studies on the frogs of the genus Leptodactylus (Amphibia: Leptodactylidae). 6. Biosystematics of the melanonotus group. Contributions in Science, 191: 1-48.

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Heyer, W. R. 1970. Gliding speeds of Ptychozoon lionatum (Reptilia: Gekkonidae) and Chrysopelea ornata (Reptilia: Colubridae). Herpetologica, 26: 317-319.

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Heyer, W. R. 1970. Studies on the genus Leptodactylus (Amphibia: Leptodactylidae). 2. Diagnosis and distribution of the Leptodactylus of Costa Rica. Revista de Biologia Tropical, 16(2): 171-205.

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Heyer, W. R. 1971. Mating calls of some frogs from Thailand. Fieldiana Zoology, 58: 61-82.

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Heyer, W. R. 1971. Descriptions of some tadpoles from Thailand. Fieldiana Zoology, 58: 83-91.

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Heyer, W. R., and Peters, J. A. 1971. The frog genus Leptodactylus in Ecuador. Proceedings of the Biological Society of Washington, 84: 163-170.

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Heyer, W. R. 1971. The tadpole of Rana pileata. Herpetological Review, 3(5): 95.

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Heyer, W. R. 1971. Leptodactylus labialis. Catalogue of American Amphibians and Reptiles, 104: 1-3.

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Heyer, W. R. 1972. A new limbless skink (Reptilia: Scincidae) from Thailand with comments on the generic status of the limbless skinks of Southeast Asia. Fieldiana Zoology, 58: 109-129.

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Heyer, W. R. 1972. The status of Leptodactylus pumilio Boulenger (Amphibia, Leptodactylidae) and the description of a new species of Leptodactylus from Ecuador. Contributions in Science, 231: 1-8.

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Heyer, W. R. 1973. Systematics of the marmoratus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae). Contributions in Science, 251: 1-50.

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Heyer, W. R. 1973. Ecological interactions of frog larvae at a seasonal tropical location in Thailand. Journal of Herpetology, 7(4): 337-361.

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Heyer, W. R., and Bellin, M. S. 1973. Ecological notes on five sympatric Leptodactylus (Amphibia, Leptodactylidae) from Ecuador. Herpetologica, 29(1): 66-72.

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Heyer, W. R., and Berven, K. A. 1973. Species diversities of herpetofaunal samples from similar microhabitats at two tropical sites. Ecology, 54(3): 642-645.

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Heyer, W. R. 1974. Niche measurements of frog larvae from a seasonal tropical location in Thailand. Ecology, 55(3): 651-656.

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Heyer, W. R. 1974. Relationships of the marmoratus species group (Amphibia, Leptodactylidae) within the subfamily Leptodactylinae. Contributions in Science, 253: 1-45.

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Heyer, W. R. 1974. Vanzolinius, a new genus proposed for Leptodactylus discodactylus (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 87(11): 81-90.

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Heyer, W. R., and Diment, J. M. 1974. The karyotype of Vanzolinius discodactylus and comments on usefulness of karyotypes in determining relationships in the Leptodactylus-complex (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 87(29): 327-336.

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McDiarmid, R. W., Heyer, W. R., and Weigmann, D. L. 1974. Tadpoles, predation and pond size in the tropics. Herpetological Review, 5(3): 76.

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Heyer, W. R. 1975. Adenomera lutzi (Amphibia: Leptodactylidae), a new species of frog from Guyana. Proceedings of the Biological Society of Washington, 88(28): 315-318.

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Heyer, W. R., McDiarmid, R. W., and Weigmann, D. L. 1975. Tadpoles, predation and pond habitats in the tropics. Biotropica, 7(2): 100-111.

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Heyer, W. R. 1975. A preliminary analysis of the intergeneric relationships of the frog family Leptodactylidae. Smithsonian Contributions to Zoology, 199: 1-55.

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Pyburn, W. F., and Heyer, W. R. 1975. Identity and call of the frog, Leptodactylus stenodema. Copeia, 3: 585-587.

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Heyer, W. R. 1976. Notes on the frog fauna of the Amazon Basin. Acta Amazonica, 6(3): 369-378.

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Heyer, W. R. 1976. Studies in larval amphibian habitat partitioning. Smithsonian Contributions to Zoology, 242: 1-27.

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Heyer, W. R. 1976. [Review] "Evolution of the Desert Biota." Copeia, 3: 614-615.

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Heyer, W. R. 1976. The presumed tadpole of Paratelmatobius lutzi (Amphibia, Leptodactylidae). Papeis Avulsos de Zoologia, 30(10): 133-135.

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Heyer, W. R., and Liem, D. S. 1976. Analysis of the intergeneric relationships of the Australian frog family Myobatrachidae. Smithsonian Contributions to Zoology, 233: 1-29.

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Heyer, W. R., and Muedeking, M. H. 1976. Notes on tadpoles as prey for naiads and turtles. Journal of Washington Academy of Sciences, 66(4): 235-239.

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Muedeking, M. H., and Heyer, W. R. 1976. Descriptions of eggs and reproductive patterns of Leptodactylus pentadactylus (Amphibia: Leptodactylidae). Herpetologica, 32: 137-139.

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Straughan, I. R., and Heyer, W. R. 1976. A functional analysis of the mating calls of the Neotropical frog genera of the Leptodactylus complex (Amphibia, Leptodactylidae). Papeis Avulsos de Zoologia, 29(23): 221-245.

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Heyer, W. R. 1977. A discriminant function analysis of the frogs of the genus Adenomera (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 89(51): 581-592.

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Heyer, W. R. 1977. Taxonomic notes on frogs from the Madeira and Purus Rivers, Brasil. Papeis Avulsos de Zoologia, 31(8): 141-162.

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Heyer, W. R. 1977. Foam nest construction in the leptodactylid frogs Leptodactylus pentadactylus and Physalaemus pustulosus (Amphibia, Anura, Leptodactylidae). Journal of Herpetology, 11(2): 225-228.

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Heyer, W. R. 1978. Systematics of the fuscus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae). National History Museum of Los Angeles County, Science Bulletin, 8(29): 1-85.

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Heyer, W. R. 1978. Variation in members of the Centrolenella eurygnatha complex (Amphibia: Centrolenidae) from Serra do Mar and Serra da Mantiqueira, Brasil. Papeis Avulsos de Zoologia, 32(2): 15-33.

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Heyer, W. R. 1978. [Review] "A Biogeographical Analysis of the Chihuahuan Desert through Its Herpetofauna." Herpetological Review, 9(2): 50-51.

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Heyer, W. R. 1979. Annual variation in larval amphibian populations within a temperate pond. Journal of the Washington Academy of Sciences, 69(2): 65-74.

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Heyer, W. R. 1979. Systematics of the pentadactylus species group of the frog genus Leptodactylus (Amphibia: Leptodactylidae). Smithsonian Contributions to Zoology, 301: 1-43.

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Heyer, W. R., and Crombie, R. I. 1979. Natural history notes on Craspedoglossa stejnegeri and Thoropa petropolitana (Amphibia: Salientia, Leptodactylidae). Journal of the Washington Academy of Sciences, 69(1): 17-20.

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Heyer, W. R., and Mello, C. C. 1979. Descriptions of the advertising calls of Cycloramphus asper and Cycloramphus dubius (Amphibia: Leptodactylidae). Papeis Avulsos de Zoologia, 32(15): 193-200.

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Heyer, W. R. 1980. The calls and taxonomic positions of Hyla giesleri and Ololygon opalina (Amphibia: Anura: Hylidae). Proceedings of the Biological Society of Washington, 93(3): 655-661.

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Maxson, L. R., Heyer, W. R., and Maxson, R. D. 1981. Phylogenetic relationships of the Brasilian leptodactylid frog genera Craspedoglossa, Cycloramphus and Zachaenus. Experientia, 37(2): 144-145.

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Drewry, G. E., Heyer, W. R., and Rand, A. S. 1982. A functional analysis of the complex call of the frog Physalaemus pustulosus. Copeia, 3: 636-645.

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Heyer, W. R. 1982. Two new species of the frog genus Hylodes from Caparao, Minas Gerais, Brasil (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 95(2): 377-385.

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Heyer, W. R., Daugherty, C. H., and Maxson, L. R. 1982. Systematic resolution of the genera of the Crinia complex (Amphibia: Anura: Myobatrachidae). Proceedings of the Biological Society of Washington, 95(3): 423-427.

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Heyer, W. R., and Maxson, L. R. 1982. Distributions, relationships, and zoogeography of lowland frogs: The Leptodactylus complex in South America, with special reference to Amazonia. In: Prance, G. T. (ed.), Biological diversification in the tropics. New York: Columbia University Press, pp. 375-388.

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Heyer, W. R., and Maxson, L. R. 1982. Neotropical frog biogeography: Paradigms and problems. American Zoologist, 22(2): 397-410.

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Maxson, L. R., and Heyer, W. R. 1982. Leptodactylid frogs and the Brasilian Shield: an old and continuing adaptive relationship. Biotropica, 14(1): 10-15.

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Crombie, R. I., and Heyer, W. R. 1983. Leptodactylus longirostris (Anura: Leptodactylidae): Advertisement call, tadpole, ecological and distributional notes. Revista Brasileira de Biologia, 43(3): 291-296.

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Heyer, W. R. 1983. Variation and systematics of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae). Arquivos de Zoologia, 30(4): 235-339.

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Heyer, W. R. 1983. Clarification of the names Rana mystacea Spix, 1824, Leptodactylus amazonicus Heyer, 1978 and a description of a new species, Leptodactylus spixi (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 96(2): 270-272.

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Heyer, W. R. 1983. Notes on the frog genus Cycloramphus (Amphibia: Leptodactylidae), with descriptions of two new species. Proceedings of the Biological Society of Washington, 96(3): 548-559.

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Heyer, W. R., and Maxson, L. R. 1983. Relationships, zoogeography, and speciation mechanisms of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae). Arquivos de Zoologia, 30(5): 341-373.

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Heyer, W. R., and Pyburn, W. F. 1983. Leptodactylus riveroi, a new frog species from Amazonia, South America (Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 96(3): 560-566.

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Wassersug, R. J., and Heyer, W. R. 1983. Morphological correlates of subaerial existence in leptodactylid tadpoles associated with flowing water. Canadian Journal of Zoology, 61(4): 761-769.

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Heyer, W. R. 1984. The systematic status of Adenomera griseigularis Henle, with comments on systematic problems in the genus Adenomera (Amphibia: Leptodactylidae). Amphibia-Reptilia, 5(2): 97-100.

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Heyer, W. R. 1984. Variation, systematics, and zoogeography of Eleutherodactylus guentheri and closely related species (Amphibia: Anura: Leptodactylidae). Smithsonian Contributions to Zoology, 402: 1-42.

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Heyer, W. R. 1984. [Review] Advances in Herpetology and Evolutionary Biology: Essays in Honor of Ernest E. Williams. Rhodin, Anders G. J., and Miyata, K. Science, 223(4632): 162.

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Heyer, W. R. 1985. Taxonomic and natural history notes on frogs of the genus Centrolenella (Amphibia: Centrolenidae) from southeastern Brasil and adjacent Argentina. Papeis Avulsos de Zoologia, 36(1): 1-21.

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Heyer, W. R. 1985. New species of frogs from Boraceia, Sao Paulo, Brazil. Proceedings of the Biological Society of Washington, 98(3): 657-671.

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Vanzolini, P. E., and Heyer, W. R. 1985. The American herpetofauna and the interchange. Topics in Geobiology, 4: 475-487.

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Heyer, W. R., and Cocroft, R. B. 1986. Descriptions of two new species of Hylodes from the Atlantic forests of Brazil (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 99(1): 100-109.

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Cocroft, R. B., and Heyer, W. R. 1988. Notes on the frog genus Thoropa (Amphibia: Leptodactylidae) with a description of a new species (Thoropa saxatilis). Proceedings of the Biological Society of Washington, 101(1): 209-220.

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Heyer, W. R. 1988. A notable collection of Cycloramphus (Amphibia, Leptodactylidae) from Bahia, Brazil, with a description of a new species (Cycloramphus migueli). Proceedings of the Biological Society of Washington, 101(1): 151-154.

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Heyer, W. R. 1988. On frog distribution patterns east of the Andes. In: Vanzolini, P. E., and Heyer, W. R. (eds.), Proceedings of a Workshop on Neotropical Distribution Patterns Held 12-16 January 1987. Rio de Janeiro: Academia Brasileira de Ciencias, pp. 245-273.

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Heyer, W. R., Rand, A. S., Cruz, C. A. G., and Peixoto, O. L. 1988. Decimations, extinctions, and colonizations of frog populations in southeast Brazil and their evolutionary implications. Biotropica, 20(3): 230-235.

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Heyer, W. R., and Vanzolini, P. E. 1988. Introduction and summary of a workshop on Neotropical distribution patterns held 12-16 January 1987. In: Vanzolini, P. E., and Heyer, W. R. (eds.), Proceedings of a Workshop on Neotropical Distribution Patterns Held 12-16 January 1987. Rio de Janeiro: Academia Brasileira de Ciencias, pp. 7-18.

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Maxson, L. R., and Heyer, W. R. 1988. Molecular systematics of the frog genus Leptodactylus (Amphibia: Leptodactylidae). Fieldiana Zoology, 41: 1-13.

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Wassersug, R. J., and Heyer, W. R. 1988. A survey of internal oral features of leptodactyloid larvae (Amphibia: Anura). Smithsonian Contributions to Zoology, 457: 1-99.

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Heyer, W. R., and Wolf, A. J. 1989. Physalaemus crombiei (Amphibia, Leptodactylidae), a new frog species from Espirito Santo, Brazil with comments on the Physalaemus-signifer group. Proceedings of the Biological Society of Washington, 102(2): 500-506.

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Heyer, W. R., Rand, A. S., Cruz, C. A. G., Peixoto, O. L., and Nelson, C. E. 1990. Frogs of Boraceia. Arquivos de Zoologia, 31(4): 231-410.

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Heyer, W. R., and Hardy, L. M. 1991. A new species of frog of the Eleutherodactylus lacrimosus assembly from Amazonia, South America (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 104(3): 436-447.

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Dubois, A., and Heyer, W. R. 1992. Leptodactylus labialis, the valid name for the American Whitelipped Frog (Amphibia: Leptodactylidae). Copeia, 2: 584-585.

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Heyer, W. R., and Perez, L. G. 1993. Description of the advertisement call and resolution of the systematic status of Leptodactylus gracilis delattini Muller, 1968 (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 106(1): 51-56.

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Heyer, W. R., Donnelly, M. A., McDiarmid, R. W., Hayek, Lee-Ann C., and Foster, M. S. 1994. Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, Washington and London. 364 pages.

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Heyer, W. R., Donnelly, M. A., McDiarmid, R. W., Hayek, Lee-Ann C., and Foster, M. S. 1994. Medicion y Monitoreo de la Diversidad Biologica: Metodos estandarizados para Anfibios. Editorial Universitaria de la Patagonia, 349 pages.

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Heyer, W. R. 1994. Variation within the Leptodactylus podicipinus-wagneri complex of frogs (Amphibia: Leptodactylidae). Smithsonian Contributions to Zoology, 546: 1-124.

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Heyer, W. R. 1994. Hyla benitezi (Amphibia: Anura: Hylidae): First record for Brazil and its biogeographical significance. Journal of Herpetology, 28(4): 497-499.

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Cardoso, A. J., and Heyer, W. R. 1995. Advertisement, aggressive, and possible seismic signals of the frog Leptodactylus syphax (Amphibia, Leptodactylidae). Alytes, 13(2): 67-76.

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Heyer, W. R. 1995. South American rocky habitat Leptodactylus (Amphibia: Anura: Leptodactylidae) with description of two new species. Proceedings of the Biological Society of Washington, 108(4): 695-716.

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Heyer, W. R., and Gascon, C. 1995. Collection notes and call characteristics for Ischnocnema quixensis and Phyzelaphryne miriamae (Amphibia: Anura; Leptodactylidae). Journal of Herpetology, 29(2): 304-307.

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Heyer, W. R., and Morales, V. R. 1995. The advertisement call of the leptodactylid frog Leptodactylus griseigularis. Amphibia-Reptilia, 16(1): 91-92.

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Heyer, W. R. 1996. Vanishing amphibians: a real problem? World Conservation. 1(96): 10-11.

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Heyer, W. R., de Sa, R. O., McCranie, J. R., and Wilson, L. D. 1996. Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae): Natural history notes, advertisement call, and relationships. Herpetological Natural History, 4(2): 169-174.

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Heyer, W. R., Garcia Lopez, J. M., and Cardoso, A. J. 1996. Advertisement call variation in the Leptodactylus mystaceus species complex (Amphibia: Leptodactylidae) with a description of a new sibling species. Amphibia-Reptilia, 17(1): 7-31.

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Halliday, T. R., and Heyer, W. R. 1997. The case of the vanishing frogs. Technology Review, 100(4): 56-63.

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Heyer, W. R. 1997. Geographic variation in the frog genus Vanzolinius (Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 110(3): 338-365.

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Savage, J. M., and Heyer, W. R. 1997. Digital webbing formulae for anurans: a refinement. Herpetological Review, 28(3): 131.

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Heyer, W. R. 1998. The relationships of Leptodactylus diedrus (Anura, Leptodactylidae). Alytes, 16(1-2): 1-24.

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Kress, W. J., Heyer, W. R., Acevedo-Rodriguez, P., Coddington, J. A., Cole, D., Erwin, T. L., Meggers, B. J., Pogue, M. G., Thorington, R. W., Jr., Vari, R. P., Weitzman, Marilyn J., and Weitzman, S. H. 1998. Amazonian biodiversity: Assessing conservation priorities with taxonomic data. Biodiversity and Conservation, 7(12): 1577-1587.

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Heyer, W. R., Coddington, Jonathan A., Kress, W. John, Acevedo-Rodriguez, Pedro, Cole, Daniel G., Erwin, Terry L., Meggers, Betty J., Pogue, Michael G., Thorington, R. W., Jr., Vari, R. P., Weitzman, M. J., and Weitzman, S. H. 1999. Amazonian biotic data and conservation decisions. Ciencia e Cultura, 51(5-6): 372-385.

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Heyer, W. R., and Munoz, A. M. 1999. Validation of Eleutherodactylus crepitans Bokermann, 1965, notes on the types and type locality of Telatrema heterodactylum Miranda-Ribeiro, 1937, and description of a new species of Eleutherodactylus from Mato Grosso, Brazil (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 112(1): 1-18.

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Heyer, W. R. 2000. The lectotype of Cystignathus hylodes Reinhardt and Lutken, 1862. Journal of Herpetology, 34(1): 150-153.

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Heyer, W. R., and Morato de Carvalho, C. 2000. Calls and calling behavior of the frog Leptodactylus natalensis (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 113(1): 284-290.

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Heyer, W. R., and Morato de Carvalho, C. 2000. The enigmatic advertisement call of Eleutherodactylus ramagii (Amphibia: Anura: Leptodactylidae). Amphibia-Reptilia, 21(1): 117-121.

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Heyer, W. R., and Thompson, A. S. 2000. Leptodacylus rugosus Noble. Catalogue of American Amphibians and Reptiles. 708: 1-5.

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Heyer, W. R. 2000. Amphibian declines and extinctions: The situation in Latin America. Ecologia Latinoamericana. 47-55.

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Carey, C., Heyer, W. R., Wilkinson, J., Alford, R. A., Arntzen, J. W., Halliday, T., Hungerford, L., Lips, K. R., Middleton, E. M., Orchard, S. A., and Rand, A. S. 2001. Amphibian declines and environmental change: Use of remote-sensing data to identify environmental correlates. Conservation Biology, 15(4): 903-913.

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Hayek, Lee-Ann C., Heyer, W. R., and Gascon, C. 2001. Frog morphometrics: A cautionary tale. Alytes, 18(3-4): 153-177.

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Heyer, W. R., and Heyer, M. M. 2001. Leptodactylus lithonaetes Heyer. Catalogue of American Amphibians and Reptiles, 723: 1-3.

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Wynn, Addison H., and Heyer, W. R. 2001. Do geographically widespread species of tropical amphibians exist? An estimate of genetic relatedness within the Neotropical frog Leptodactylus fuscus (Schneider 1799) (Anura Leptodactylidae). Tropical Zoology, 14: 255-285.

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Heyer, W. R. 2002. Leptodactylus fragilis, the valid name for the Middle American and northern South American white-lipped frog (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 115(2): 321-322.

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Heyer, W. R., and Caramaschi, U. 2002. Comment on the proposed conservation of the specific name of Leptodactylus chaquensis Cei, 1950 (Amphibia, Anura). Bulletin of Zoological Nomenclature, 59(1): 44-45.

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Heyer, W. R., de Sa, R. O., and Muller, S. 2002. Leptodactylus silvanimbus McCranie, Wilson, and Porras. Catalogue of American Amphibians and Reptiles, 743: 1-3.

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Heyer, W. R., and Heyer, M. M. 2002. Leptodactylus elenae Heyer. Catalogue of American Amphibians and Reptiles, 742: 1-5.

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Heyer, M. M., Heyer, W. R., Spear, S., and de Sa, R. O. 2003. Leptodactylus mystacinus (Burmeister) Mustached Frog. Catalogue of American Amphibians and Reptiles. 767: 1-11.

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Heyer, W. R., and Junca, Flora A. 2003. Leptodactylus caatingae, a new species of frog from eastern Brazil (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 116(2): 317-329.

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Heyer, W. R., and Reid, Yana R. 2003. Does advertisement call variation coincide with genetic variation in the genetically diverse frog taxon currently known as Leptodactylus fuscus (Amphibia: Leptodactylidae)? Anais da Academia Brasileira de Ciencias, 75(1): 39-54.

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Heyer, W. R. 2004. Paulo Emilio Vanzolini. Copeia, 2004(1): 184-189.

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Heyer, W. R., and Heyer, M. M. 2004. Leptodactylus furnarius Sazima and Bokermann cerrado oven frog. Catalogue of American Amphibians and Reptiles, 785: 1-5.

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Brandao, R. A., and Heyer, W. R. 2005. The complex calls of Leptodactylus pustulatus (Amphibia, Anura, Leptodactylidae). Amphibia-Reptilia, 26(4): 566-570.

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Caramaschi, U., de Sa, R. O., and Heyer, W. R. 2005. Common names for the frog genus Leptodactylus (Amphibia, Anura, Leptodactylidae). Herpetological Review, 36(2): 119-120.

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de Sa, R. O., Heyer, W. R., and Camargo, A. 2005. A phylogenetic analysis of Vanzolinius Heyer, 1974 (Amphibia, Anura, Leptodactylidae): Taxonomic and life history implications. Arquivos do Museu Nacional, 63(4): 707-726.

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de Sa, R. O., Heyer, W. R., and Camargo, A. 2005. Are Leptodactylus didymus and L. mystaceus phylogenetically sibling species (Amphibia, Anura, Leptodactylidae)? Herpetologia Petropolitana, Russian Journal of Herpetology, 12: 90-92.

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Hayek, Lee-Ann C., and Heyer, W. R. 2005. Determining sexual dimorphism in frog measurement data: Integration of statistical significance, measurement error, effect size and biological significance. Anais da Academia Brasileira de Ciencias, 77(1): 45-76.

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Heyer, W. R., de Sa, R. O., and Muller, S. 2005. On the enigmatic distribution of the Honduran endemic Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae). In: Donnelly, M. A., Crother, B. I., Guyer, C., Wake, M. H., and White, M. E. (eds.), Ecology and Evolution in the Tropics: A Herpetological Perspective. Chicago: University of Chicago Press, pp. 81-101.

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Heyer, W. R., and Murphy, J. B. 2005. Declining Amphibian Populations Task Force. In: Lannoo, Michael (ed.), Amphibian Declines: The Conservation Status of United States Species. Berkeley, CA: University of California Press, pp. 17-21.

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Heyer, W. R. 2005. Leptodactylus fragilis Brocchi, 1877. White-lipped frog. In: Lannoo, M. (ed.), Amphibian Declines: The Conservation Status of United States Species. Berkeley, CA: University of California Press, pp. 500-501.

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Heyer, W. R., and Crombie, R. I. 2005. Leptodactylus lauramiriamae, a distinctive new species of frog (Amphibia: Anura: Leptodactylidae) from Rondonia, Brazil. Proceedings of the Biological Society of Washington, 118(3): 590-595.

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Heyer, W. R. 2005. Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, northern South America, and Amazonia. Arquivos de Zoologia, 37(3): 269-348.

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Heyer, W. R., de Sa, R. O., and Rettig, A. 2005. Sibling species, advertisement calls, and reproductive isolation in frogs of the Leptodactylus pentadactylus species cluster (Amphibia, Leptodactylidae). Herpetologia Petropolitana, Russian Journal of Herpetology, 12: 35-39.

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Camargo, Arley, de Sa, R. O., and Heyer, W. R. 2006. Phylogenetic analyses of mtDNA sequences reveal three cryptic lineages in the widespread neotropical frog Leptodactylus fuscus (Schneider, 1799) (Anura, Leptodactylidae). Biological Journal of the Linnean Society, 87(2): 325-341.

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Heyer, W. R. 2006. Obituary: In celebration of A. Stanley Rand (1932-2005). Phyllomedusa, 5(1): 8-10.

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Heyer, W. R., and Heyer, M. M. 2006. Leptodactylus knudseni Heyer, Knudsen's thin-toed frog. Catalogue of American Amphibians and Reptiles, 807: 1-12.

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Heyer, W. R., and Heyer, M. M. 2006. Leptodactylus natalensis A. Lutz, bubbling frog. Catalogue of American Amphibians and Reptiles, 808: 1-5.

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Heyer, M. M., Heyer, W. R., and de Sa, R. O. 2006. Leptodactylus fragilis (Brocchi) White-lipped Thin-toed Frog. Catalogue of American Amphibians and Reptiles, 830: 1-26.

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Heyer, W. R., and Scott, N. J., Jr. 2006. The advertisement call of Leptodactylus laticeps (Amphibia, Anura, Leptodactylidae): predatory aural luring? Herpetological Natural History, 9(2): 189-194.

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Heyer, W. R., Caramaschi, U., and de Sa, R. O. 2006. Case 3323. Rana ocellata Linnaeus, 1758 (currently Leptodactylus ocellatus; Amphibia, Anura): proposed conservation of usage of the specific name by the designation of a neotype. Bulletin of Zoological Nomenclature, 63(3): 184-186.

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Yanek, K., Heyer, W. R., and de Sa, R. O. 2006. Genetic resolution of the enigmatic Lesser Antillean distribution of the frog Leptodactylus validus (Anura, Leptodactylidae). South American Journal of Herpetology, 1(3): 192-201.

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Ponssa, M. L., and Heyer, W. R. 2007. Osteological characterization of four putative species of the genus Adenomera (Anura: Leptodactylidae), with comments on intra- and interspecific variation. Zootaxa, 1403: 37-54.

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Bolanos, F., Castro, F., De la Riva, I., Grant, T., Hedges, B., Heyer, W. R., Ibanez D., Roberto, La Marca, E., Lavilla, E., Leite Silvano, D., Lotters, S., Parra Olea, G., Reichle, S., Reynolds, Robert P., Rodriguez, L., Santos Barrera, G., Scott, N., Ubeda, C., Veloso, A., Wilkinson, M., and Young, B. 2008. Amphibians of the Neotropical Realm. In: Stuart, S. N., Hoffmann, M., Chanson, J. S., Cox, N. A., Berridge, R. J., Ramani, P., and Young, B. E. (eds.), Threatened Amphibians of the World. Barcelona: Lynx Edicions, pp. 92-105.

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Heyer, W. R., Heyer, M. M., and de Sa, R. O. 2008. Leptodactylus cunicularius Sazima and Bokermann, rabbit-burrow frog. Catalogue of American Amphibians and Reptiles, 845: 1-5.

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Heyer, W. R. 2008. Can long-established scientific societies survive the digital age? Journal of the Washington Academy of Sciences, 94(2): 17-32.

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Camargo, Arley, Heyer, W. R., and de Sa, R. O. 2009. Phylogeography of the frog Leptodactylus validus (Amphibia: Anura): Patterns and timing of colonization events in the Lesser Antilles. Molecular Phylogenetics and Evolution, 53: 571-579.

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Fernandez, M., Cole, D. G., Heyer, W. R., Reichle, Steffen, and de Sa, R. O. 2009. Predicting Leptodactylus (Amphibia, Anura, Leptodactylidae) distributions: Broad-ranging versus patchily distributed species using a presence-only environmental niche modeling technique. South American Journal of Herpetology, 4(2): 103-116.

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Heyer, M. H., Heyer, W. R. and de Sa, R. O. 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Amphibia, Anura, Leptodactylidae). Volume 1. References. Smithsonian Herpetological Information Service, 1: 1-677.; Heyer, M. H., Heyer, W. R., and de Sa, R. O. 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Amphibia, Anura, Leptodactylidae). Volume 2. Indices. Smithsonian Herpetological Information Service, 2: 1-167. (electronic records)

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Heyer, W. R., and Barrio-Amoros, C. L. 2009. The advertisement calls of two sympatric frogs, Leptodactylus lithonaetes (Amphibia: Anura: Leptodactylidae) and Pristimantis vilarsi (Amphibia: Anura: Strabomantidae). Proceedings of the Biological Society of Washington, 122(3): 282-291.

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Heyer, W. R., and Giaretta, A. A. 2009. Advertisement calls, notes on natural history, and distribution of Leptodactylus chaquensis (Amphibia: Anura: Leptodactylidae) in Brasil. Proceedings of the Biological Society of Washington, 122(3): 292-305.

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Furness, A. I., McDiarmid, R. W., Heyer, W. R., and Zug, G. R. 2010. Oviduct modifications in foam-nesting frogs, with emphasis on the genus Leptodactylus (Amphibia, Leptodactylidae). South American Journal of Herpetology, 5(1): 13-29.

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Heyer, W. R., Heyer, M. M., and de Sa, R. O. 2010. Leptodactylus savagei Heyer Savage's thin-toed frog. Catalogue of American Amphibians and Reptiles, 867: 1-19.

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Heyer, W. R., Heyer, M. M., and de Sa, R. O. 2010. Leptodactylus syphax Bokermann Burgundy thin-toed frog. Catalogue of American Amphibians and Reptiles, 868: 1-9.

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Lavilla, E. O., Langone, J. A., Caramaschi, U., Heyer, W. R., and de Sa, R. O. 2010. The identification of Rana ocellata Linnaeus, 1758. Nomenclatural impact on the species currently known as Leptodactylus ocellatus (Leptodactylidae) and Osteopilus brunneus (Gosse, 1851) (Hylidae). Zootaxa, 2346: 1-16.

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Ponssa, M. L., Heyer, W. R., and de Sa, R. O. 2010. Phylogeny of the genus Leptodactylus (Anura, Leptodactylidae) based on characters of morphology and mode of life. Cladistics, 26(2): 222.

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Heyer, W. R., and de Sa, R. O. 2011. Variation, Systematics, and Relationships of the Leptodactylus bolivianus Complex (Amphibia: Anura: Leptodactylidae). Smithsonian Contributions to Zoology, 635: 1-58.

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Heyer, M. M., Heyer, W. R., and de Sa, R. O. 2011. Leptodactylus pentadactylus (Laurenti) smoky jungle frog. Catalogue of American Amphibians and Reptiles, 887: 1-48.

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Heyer, W. R., and Heyer, M. M. 2012. Systematics, distribution, and bibliography of the frog Leptodactylus validus Garman, 1888 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 125(3): 276-294.

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Heyer, W. R. 2013. Paulo Emilio Vanzolini, 25 April 1924-30 April 2013. Herpetological Review, 44(3): 362-364.

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Heyer, W. R., and Heyer, M. M. 2013. Systematics, distribution, and bibliography of the frog Leptodactylus insularum Barbour, 1906 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 126(3): 204-233.

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Heyer, W. R. 2014. Morphological analyses of frogs of the Leptodactylus latrans complex (Amphibia, Leptodactylidae) from selected localities in South America. Proceedings of the Biological Society of Washington, 126(4): 369-378.

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