Accession 17-132

Adams, Dean C., 2004-2005

Ahlander, Erik, 2003, 2005-2006, 2010-2011, 2015

Allison, Allen, 2006

Almeida, Antonio de Padua, 2006, 2010

Almendariz, Ana, 2012

Altig, Ronald G., 2012

Amaro-Ghilardi, Renata Cecilia, 2002

Ananjeva, Natalia, 2002

Anderson, Karen, 1981, 1995

Annunziata, Bruno Barcellos, 2007

Anstis, Marion, 2005-2006, 2008

Araujo, Olivia Gabriela, 2009, 2013

Arntzen, J. W., 2004, 2007

Arruda, Mauricio Papa de, 2006

Arzabe, Cristina, 2003

Avila Pires, Teresa, 2004, 2009

Ayarzaguena, Jose, 1990

Badillo, Alberto Fernandez, 1990

Baeta, Delio, 2010

Ballentine, Barbara, 2009

Barg, Monica, 2001

Barrio Amoros, Cesar Luis, 2011-2013

Basso, Nestor, 2009

Bauer, Aaron M., 2008-2009

Berneck, Bianca, 2008

Biancardi, Carlo, 2009

Bickford, David, 2010

Bilate, Marcos, 2008

Bionda, Clarisa de Lourdes, 2009

Bisbal, Francisco, 1990

Bogart, Jim, 2006

Bohme, Wolfgang, 2003

Boistel, Renaud, 2001, 2005

Briggs, Jeffrey, 2005

Bronikowski, Ed, 2008

Bryan, Ann Wascher, 2011

Burge, Russell, 2004

Bustamante, Martin R., 2008

Caldwell, Janalee P., 2004, 2006, 2008-2010

Cannatella, David, 2002-2003, 2006-2007

Caramaschi, Ulisses, 1997, 2000-2004, 2006-2009

Cardoso, Manoela, 2010

Cardozo, Martin, 2004-2005

Carstensen, Daniel Wisbech, 2006

Carvalho, Celso Morato de, 1998, 2008-2012

Carvalho, Thiago Ribeiro de, 2012, 2015

Cascon, Paulo, 2008

Cei, Jose M., 2004

Chestnut, Tara, 2009

Cisneros-Heredia, Diego F., 2006

Cocroft, Rex, 2006

Coddington, Jonathan, 2012

Cole, Charles J., 2003, 2007-2010

Colli, Guarino R., 2003, 2009-2010

Cordova Santa Gadea, Jesus H., 2006

Costa, Henrique Caldeira, 2009, 2012

Cox, Neil, 2006-2007

Crawford, Andrew J., 2006, 2010

Crump, Martha, 2005

Cruz, Carlos Alberto, 2012-2013

Curcio, Felipe Franco, 2009

Czapracki, Allison, 2009

David, Patrick, 2004

de Queiroz, Kevin, 2007

de Ruiter, Ronald, 2009

de Sa, Rafael, 2008-2015

Dias, Iuri Ribeiro, 2008, 2011

Dixon, James R., 1998, 2000, 2002, 2005, 2007

Dodd, Kenneth, 2009

Donnelly, Maureen A., 2009

Drewery, George

Duarte B. Filho, Jose, 2009

Dundee, Harold, 2009

Erdtmann, Luciana, 2008

Ernst, Carl H., 2013

Espanola, Asociacion Herpetologica, 2008

Eterovick, Paula, 2004-2005, 2007

Fabrezi, Marissa, 2007

Faivovich, Julian, 2002, 2010

Feio, Renato, 2008

Felsenstein, Joseph, 1976

Fong G., Ansel, 2007

Ford, Linda S., 1979-1980, 1988, 1990, 1992, 1998

Forlani, Mauricio da Cruz, 2009, 2012-2013

Fouquet, Antoine, 2009-2010, 2012

Frost, Darrel, 1997, 2005-2006, 2008, 2012

Franklin, Carl J., 2010

Freitas, Joseana, 2011

Gagliardi Urrutia, Giuseppe, 2010

Garcia, Paulo C. A., 2011, 2013

Garda, Adrian Antonio, 2009

Gascon, Claude, 2011

Ghilardi, Renata Cecilia Amaro, 2004

Giaretta, Ariovaldo Antonio, 2006-2008, 2011

Giasson, Luis Menta, 2007

Gibula, Jeff, 2008

Good, Tatjana, 2007

Gordo, Marcelo, 2007

Gorzula, Stefan, 1990

Grant, Taran, 2006-2007, 2009, 2013

Green, David E., 2001

Guayasamin, Juan Manuel, 2005

Guerreiro Cordova, Manuel Alejandro, 2008

Haddad, Celio F. B., 1993, 2003-2004

Hanken, James, 2006

Hansen, Robert W., 2013

Hartsell, Traci D., 2009

Hawley, Tanya, 2007

Hayek, Lee-Ann, 2001

Henderson, Robert, 2006

Henle, Klaus, 2003

Hepp, Fabio Silva Fernandes dos Santos, 2012

Hero, Jean-Marc, 2007, 2010

Herrera Villalobos, Alvaro, 2006-2007

Hibbitts, Toby, 2005, 2007, 2010

Hicks, Tracy

Hoogmoed, Marinus, 2011-2012, 2014

Hoyer, Richard F., 2004

Hurme, Kristiina J., 2004, 2012-2014

Ibanez, Roberto, 2005, 2007

Inga, Jorge Joel, 2009

Inger, Robert F., 2010

Jackson, Kate, 2009

Jayne, Bruce C., 1988

Johnson, G. David, 2007

Johnson, Susan, 2003

Joventino Roberto, Igor, 2006, 2014

Jowers, Michael, 2003

Junca, Flora Acuna, 2000-2002, 2004-2005, 2011

Jungfer, Karl-Heinz, 2009

Kaiser, Hinrich, 2005

Kaplan, Moises, 2004

Kattan, Gustavo, 2008

King, Jay, 2012

Kizirian, David, 2007

Klaczko, Julia, 2005

Koehler, Gunther, 2011-2012

Kok, Philippe, 2006

Kokubum, Marcelo N. de C., 2013

Kozak, Kenneth H., 2007

Kretzschmar, Sonia, 2003, 2010

Kullander, Sven O., 2003

Kwet, Axel, 2004-2005, 2007

Lacerda, Joao Victor A. de, 2009

La Marca, Enrique, 2011

Langstroth, Robert, 2010

Lannoo, Michael J., 2002

Lazell, Skip, 2005

Lee, David, 2008

Leite, Felipe S. F., 2008

Lips, Karen, 2008-2009

Lobo, Fernando, 2007

Loebmann, Daniel, 2006

Loman, Jon, 2009

Losos, Jonathan B., 2000

Lutz, Adolfo

Luhring, Thomas M., 2007

Lynch, John D., 2004, 2013, 2015

MacCulloch, Ross, 2006

Magnusson, William, 2008

Manzanilla, Jesus, 2006

Marquez, Rafael, 2011

Marx, Hymen, 2007

Maxson, Linda E. R., 1979-1980, 2007

May, Heather A., 2008

Mayer, Gregory C., 2006

McCracken, Shawn F., 2009

McCranie, James R., 2012

McDiarmid, Roy W., 2008

McGuire, Jimmy A., 2007

McLeod, David S., 2005

McQuoid-Greason, Meredith, 2011

Medina, Regina Gabriela, 2013

Mello, Caio Marinho, 2014

Mello, Carolina, 2009, 2011

Melo, Paulo, 2011

Mendelson, Joseph R., III, 2008

Mendoza Roldan, Juan Salvador, 2014

Meneghelli, Diego, 2011-2012

Mijarres, Abraham, 1999, 2011

Minter, Leslie, 2006

Mitchell, Joseph C., 2007

Moen, Daniel S., 2011

Moler, Paul, 2002

Molur, Sanjay, 1999

Morgan, Gary S., 2009

Moro, Silvia, 1996

Munoz, Arturo, 2005

Murphy, John C., 2011

Myers, Charles W., 2011

Nascimento, Luciana Barreto, 2005, 2007, 2010

Nomura, Hitoshi, 2004

Nunes, Ivan, 2006, 2008, 2013, 2015

Ohler, Annemarie, 2009

Oliveira, Leonardo de, 2010

Orrico, Victor G. Dill, 2010

Ortega-Andrade, H. Mauricio, 2010

Ospina-Sarria, Jhon Jairo, 2012

Ouboter, Paul E., 2010

Oyamaguchi, Hilton, 2008

Padial, Jose, 2011

Paez V., Monica, 2006

Parham, James, 2007

Pase, H. A. (Joe), III, 2010

Passos, Paulo, 2004

Pauly, Gregory B., 2004, 2014

Pauwels, Olivier S. G., 2008

Peao, Carlos, 2007

Pearson, David L., 2003

Peloso, Pedro L. V., 2011

Peltzer, Paola, 2004

Peres, Juliana, 2008

Perry, Gad, 2006

Peters, Gustav, 2005

Pinedo, Jorge Joel, 2010

Pinto, Gabriel Silva, 2000, 2004

Pombal, Jose P., Jr., 2011

Ponssa, Maria Laura, 2005, 2011

Pontes, Rafael Cunha, 2010-2011

Prado, Cynthia P. A., 2005

Prado, Gustavo M., 2005

Pregill, Gregory K., 2007

Provete, Diogo Borges, 2008

Prudente, Ana, 2007-2008

Rapp, Tain , 2008

Relyea, Rick, 2009, 2011

Resetar, Alan, 2008

Ribeiro, Joana, 2014

Ribeiro, Marco Antonio, Jr., 2010

Rice, Jeff, 2007

Roberto, Igor Joventino, 2005-2006

Robertson, Jeanne M., 2005-2006

Robinson, Earle, 2006

Rocek, Zbynek, 2005

Rocha, Carlos Frederico Duarte da, 2006

Rocha, Mariane Targino, 2006, 2008-2009

Rodriguez Bayona, Lily Olga, 2010-2011

Roedel, Mark-Oliver, 2007

Roelofs, Terry, 2010

Romero, Heron, 2006

Ron, Santiago, 2005-2006, 2011

Rosado, Jose, 2011

Rossman, Doug, 2008

Samper, Cristian, 2007, 2009

Samyn, Yves, 2008-2009

Sanchez Ramirez, David, 2004-2005

Sanchez Pacheco, Santiago J., 2009

Santos, Jose W. A., 2008

Santos, Juan C., 2006

Santos, Marcus Thadeu Teixeira, 2013

Savage, Jay M., 2012

Savitzky, Alan H., 2008-2009

Sazima, Ivan, 1980, 2003, 2007, 2014

Scharlach, Cecilia, 2009

Schiesari, Luis, 2005, 2008

Schmid, Michael, 2005

Schulze, Arne, 2010

Segalla, Magno, 2009, 2011

Senaris, J. Celsa, 2011

Seripierri, Dione, 2002, 2006

Sheridan, Jennifer, 2005-2006

Silva, Helio Ricardo da, 2003, 2006, 2011

Silva-Filho, Ivan Sergio N., 2006

Silva, Monica Carolina Cardoso da, 2008-2009

Silva-Soares, Thiago, 2012-2013

Simmons, John E., 2004

Simon, Monique N., 2012-2013

Slough, Brian G., 2003

Smith, Joanna, 2011

Smith, Paul, 2011

Smythe, Ashleigh, 2008

Solano de Chacin, Haydee, 1990

Sole, Mirco, 2006-2007, 2010-2011

Souza, Sergio Marques de, 2008

Spencer, Kate, 1994

Stafford, Peter J., 2007

Starr, Christopher K., 2005

Strader, Ginger, 2009

Strussmann, Christine, 2004

Stuart, Simon N., 2006, 2008

Tarano, Zaida, 2011

Teixeira Junior, Mauro, 2011

Teixeira, Rogerio, 2005

Tol, J. van, 2007

Torres, Omar, 2008

Torres-Cervantes, Ricardo, 2005

Trueb, Linda, 2006, 2008

Uzzell, Thomas, 2009

Val, Aldoberto, 2006

Val, Francisca Carolina (Chica) do, 1991, 1994, 1997-2015

Vanzolini, Paulo Emilio, 2013

Vari, Richard, 2012

Veauvy, Mariana, 2008

Velez R., Caudia Maria, 2004

Verdade, Vanessa K., 2004-2007

Vieira, Kleber S., 2007, 2009

Vigle, Greg, 2006

Vogt, Richard, 2005-2008

Voris, Harold K., 2008

Wake, David, 2007-2008

Wake, Marvalee H., 2005, 2007

Waldez, Fabiano, 2007

Walker F. de Faria, Marina, 2012

Walley, Harlan D., 2007

Wassersug, Richard, 2006, 2009

Weigt, Lee, 2010

Williams, Jorge, 2005, 2009

Wilson, Don, 2009

Wray, Ken, 2008

Wynn, Addison, 2011, 2014

Yanez-Munoz, Mario Humberto, 2006, 2009

Zaher, Hussam, 2004-2006, 2008-2013

Zamudio, Kelly R., 2004-2006, 2012

Zimmerman, Barbara Lewis, 1991, 2001

Data for presentation at Annual Meeting of the Society of Integrative and Comparative Biology, Patras, Greece, 2002

Leptodactylus troglodytes call analysis for 1995 manuscript

Presentation: 246 years of Nomenclatural Novelties in the Genus Leptodactylus: Past, Present, Future

Brazilian tadpoles, data and illustrations

Brazilian tadpoles, references

Leptodactylus gazetteer

Smithsonian Amphibian Initiative, 2008

Texas collections trip, March 2010

National Public Radio "Talk of the Nation: Science Friday" presentation on declining amphibian problem, 5/16/1997 (audiocassette)

Leptodactylus validus manuscript and data, Proceedings of the Biological Society of Washington, 125(2): 276-294, 2012

Leptodactylus annotated bibliography, circa 1965- : A-B

Leptodactylus annotated bibliography, circa 1965- : C-E

Leptodactylus annotated bibliography, circa 1965- : F-H

Leptodactylus annotated bibliography, circa 1965- : I-lutzi

Leptodactylus annotated bibliography, circa 1965- : macroblepharus-notoaktites

Leptodactylus annotated bibliography, circa 1965- : occidentalis-pentadactylus

Leptodactylus annotated bibliography, circa 1965- : perlaevis-R

Leptodactylus annotated bibliography, circa 1965- : S-wuchereri

L. fuscus group standards

L. latrans group standards

L. pentadactylus group standards

Variation, Systematics, and Relationships of the Leptodactylus bolivianus Complex, 2011 (final draft)

Paulo Emilio Vanzolini background obituary materials and drafts, 2013

L. pentadactylus biogeography project: Statistics and data

L. insularum manuscript, Proceedings of the Biological Society of Washington, 126 (3): 204-233, 2013 (2 folders)

L. insularum bibliography, Proceedings of the Biological Society of Washington, 126 (3): 204-233, 2013

FAPESP (Sao Paulo Research Foundation) proposal, Leptodactylus at the Museu de Zoologia da Universidade de Sao Paulo (MZUSP), 2010

Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, northern South America, and Amazonia, Arquivos de Zoologia, 37(3): 269-348, 2005 - Measurement data coding

Leptodactylus pentadactylus, Catalogue of American Amphibians and Reptiles, (888): 1-47, 2011 - Correspondence, figures, etc.

Leptodactylus pentadactylus, Catalogue of American Amphibians and Reptiles, (888): 1-47, 2011 - Data, drafts, annotated references (5 folders)

L. latrans group data sheets

L. latrans complex, photographs/data - Argentina: San Juan, San Luis, Santa Fe, Santiago del Estero

L. latrans complex, photographs/data - Argentina: Tucuman

L. latrans complex, data sheets - Argentina: Buenos Aires, Chaco, Cordoba, Corrientes, Entre Rios, Formosa, Jujuy, La Pampa, Mendoza, Misiones, Salta, San Juan, San Luis, Santa Fe, Santiago del Estero, Tucuman

L. latrans complex, photographs/data - Bolivia: Beni, Chuquisaca, La Paz

L. latrans complex, photographs/data - Bolivia: Santa Cruz

L. latrans complex, photographs/data - Bolivia: Beni, Chuquisaca, La Paz, Santa Cruz

L. latrans complex, photographs/data - Brasil: Alagoas, Amapa, Amazonas, Bahia, Ceara, Distrito Federal, Espirito Santo

L. latrans complex, photographs/data - Brasil: Amazonas - Coari

L. latrans complex, photographs/data - Argentina: No locality, Distrito Buenos Aires, Chaco

L. latrans complex, photographs/data - Argentina: Cordoba, Corrientes

L. latrans complex, photographs/data - Argentina: Entre Rios, Formosa

L. latrans complex, photographs/data - Argentina: Jujuy, La Pampa, Mendoza, Misiones, Neuquen, Salta

L. latrans complex, photographs/data - Brasil: Amazonas - Coari

L. latrans complex, photographs/data - Brasil: Goias

L. latrans complex, photographs/data - Brasil: Maranhao, Mato Grosso

L. latrans complex, photographs/data - Brasil: Mato Grosso do Sul

L. latrans complex, photographs/data - Brasil: Minas Gerais

L. latrans complex, photographs/data - Brasil: Para, Paraiba, Pernambuco, Piaui, Rio de Janeiro

L. latrans complex, photographs/data - Brasil: Rio Grande do Norte, Rondonia, Roraima, Santa Catarina, Sao Paulo, Sergipe, Tocantins

L. latrans complex, data sheets - Brasil: Acre, Alagoas, Amapa, Amazonas - Coari, Bahia, Ceara, Distrito Federal, Goias

L. latrans complex, data sheets - Brasil: Maranhao, Mato Grosso, Mato Grosso do Sul, Minas Gerais, Para, Paraiba, Parana, Pernambuco, Piaui

L. latrans complex, data sheets - Brasil: Rio de Janeiro, Rio Grande do Sul, Rondonia, Roraima, Santa Catarina, Sao Paulo, Sergipe, Tocantins

L. latrans complex, photographs/data - Guyana; Venezuela: Apure, Bolivar

L. latrans complex, photographs/data - Venezuela: Guarico

L. latrans complex, data sheets - Guyana; Venezuela: Apure, Bolivar, Guarico

L. latrans complex, photographs/data - Trinidad

L. latrans complex, photographs/data - Paraguay: Alto Paraguay, Amambay, Boqueron, Canindeyu, Central, Cordillera, Itapua

L. latrans complex, photographs/data - Paraguay: Paraguari, Presidente Hayes, San Pedro

L. latrans complex, photographs/data - Uruguay: Canelones, Maldonado, Rivera, Rocha, Treinta y Tres

L. latrans complex, data sheets - Paraguay, Uruguay

L. latrans complex - Preliminary discriminant function analyses

L. latrans complex - Miscellaneous data

L. latrans complex - Analyses from digital photographs (as L. ocellatus)

L. latrans complex - Regional analyses of morphological data manuscript

L. latrans complex - Discriminant function analyses data, draft, regional samples

L. latrans complex - Manuscript, various documents

Slides of Brasilian field work sites and animals

Dissertation plates

Publication illustrations, originals (includes electronic records)

Leptodactylus validus poster presentation

"The Last Frog" National Geographic presentation (includes VHS)

Howard University presentation - "Why Study Frogs from Nature?", 1989

"Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians," online chapters

"Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians," 1994 (English)

"Medicion y Monitoreo de la Diversidad Biologica: Metodos estandarizados para Anfibios," 1994 (Spanish)

"Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians" (Russian)

Amazon presentation - Curitiba herpetology meetings

Heyer and de Sa National Science Foundation (NSF) proposal, 1996-1997

Heyer and de Sa NSF award 9815787

Heyer and de Sa NSF proposal, 2003

Heyer and de Sa NSF grant records and finances as of July 2004

Leptodactylus phylogeny data and correspondence

Leptodactylus phylogeny monograph final publication, South American Journal of Herpetology Special Issue, 9 (S1-S128), 2014

Leptodactylus phylogeny monograph drafts and data (6 folders)

Leptodactylus phylogeny frog call recordings and requests, photographs and requests

Leptodactylus phylogeny tissue data and POY runs

Leptodactylus phylogeny tree

Leptodactylus phylogeny tissues data

Leptodactylus phylogeny monograph correspondence

Cornell Laboratory of Ornithology NSF proposal for frog recordings

Leptodactylus website common names, calls, photographs

Unpublished Leptodactylus sexual dimorphism manuscript, 2015

Sexual dimorphism data analysis: Leptodactylus albilabris - Leptodactylus discodactylus

Sexual dimorphism data analysis: Leptodactylus elenae - Leptodactylus insularum

Sexual dimorphism data analysis: Leptodactylus knudseni - Leptodactylus mystacinus

Sexual dimorphism data analysis: Leptodactylus natalensis - Leptodactylus pustulatus

Sexual dimorphism data analysis: Leptodactylus rhodomerus - Leptodactylus turimiquensis

Sexual dimorphism data analysis: Leptodactylus validus - Leptodactylus wagneri

Sexual dimorphism manuscript initial drafts (includes electronic records)

Sexual dimorphism manuscript data and correspondence

Seminar presentations notes

Presentation slides: "Relationships of Leptodactylus silvanimbus"

Presentation slides: Wynn and Heyer, "Leptodactylus fuscus, one or more species"

Presentation slides: "Speciation factors in Leptodactylus fuscus, Leptodactylus pentadactylus, and Leptodactylus melanonotus groups"

Heyer manuscripts (electronic records)

Leptodactylus pdf files (electronic records)

Estuarium frog talk (electronic records)

Speciation patterns in L. pentadactylus cluster (electronic records)

Brazilian Herpetological Society Belo Horizonte presentation (electronic records)

Leptodactylus bolivianus complex presentation (electronic records)

EndNote presentation, Leptodactylus research topics, Rio de Janeiro and San Juan (electronic records)

Belem, Brazil, presentation (electronic records)

Leptodactylus systematics (electronic records)

Argentina presentation (electronic records)

7th Congress, Argentina, presentation (electronic records)

San Luis, Argentina, presentation (electronic records)

Locality records - Thailand

Locality data and correspondence - South America

Leptodactylus locality data (2 folders)

Leptodactylus locality Gazetteer

Drawings, sonograms, maps, data, and photographs for publication

Marmoratus group - locality data

Venezuelan Leptodactylus species data

Leptodactylus locality data

JAP specimens to borrow

Negatives

External morphology data

Leptodactylus - Tadpole data

Barycholos

Lithodytes

External morphology of melanonotus group

Leptodactylus - Egg and egg deposition information

Leptodactylus - Taxonomic notes and miscellaneous

Chromosome analysis

L. labialis - Analyses

L. albilabris - Analyses

Leptodactylus locality data

Fuscus group data

Mystaceus - SW complex form

Mystaceus

Fuscus group - Nomenclature correspondence

Completed data sheets

Pentadactylus group nomenclature

Leptodactylus data sheets

Leptodactylus - Breeding call information

Leptodactylid literature

Chromosome data

Heyer and Pyburn - Leptodactylus riveroi

Pattern standards - Melanonotus group

Post podicipinus-wagneri manuscript locality data

Leptodactylus - Distributional data post revisions

Cycloramphus negatives

Centrolenella negatives

Negatives for fuscus group paper

Map negatives for fuscus group paper

Proof sheet and negatives for figures associated with L. rugosus, etc., manuscript

Leptodactylus paraguayensis and L. didymus negatives

Map negative of South America

Leptodactylus type negatives

E. guentheri negatives

Podicipinus-wagneri complex negatives

Large species of Leptodactylus holotype negatives and proof sheet

Eleuth. dundeei, Rupirana cardosoi, Rondonia frog negatives

Unidentified negatives (2 folders)

Fuscus group standard sheets

Fuscus group analyses and notes for manuscript

Tepequem collection notes

Cadle Bolivian frog

Types - Senckenberg, Vienna, Munich, September 1982

Leptodactylus types A-L

Leptodactylus types M-Z

L. delattini

L. cf furnarius - Pico das Almas

L. ocellatus

L. species "elenae" from Peru

Albilabris - Data and transformations

L. bolivianus

L. bufonius

L. caatingae

L. camaquara

L. chaquensis

L. colombiensis

L. cunicularius

L. cupreus

L. didymus - Data

L. diedrus

Vanzolinius discodactylus

L. elenae

L. fallax

L. flavopictus

L. furnarius

L. fragilis

Fuscus - Data

L. jolyi

L. knudseni

L. labrosus

L. labyrinthicus

L. laticeps

L. latinasus

L. gracilis

L. griseigularis

L. insularum

L. lauramiriamae

L. leptodactyloides

L. lithonaetes

L. longirostris

L. macrosternum

L. magistris

L. marambaiae

L. melanonotus

L. myersi

L. mystaceus

L. mystacinus

L. natalensis

L. nesiotus

L. notoakfites

L. ocellatus

L. paraensis

L. pascoensis

L. pentadactylus - Amazonia

L. peritoaktites

L. petersii

L. plaumanni

L. podicipinus

L. poecilochilus

L. pustulatus

Ceara L. pustulatus

L. rhodomerus

L. rhodomystax

L. rhodonotus

L. riveroi

L. rugosus

L. sabanensis

L. savagei

L. silvanimbus

L. spixi

L. stenodema

L. syphax

L. tapiti

L. troglodytes

L. turimiquensis

L. validus

L. vastus

L. ventrimaculatus

L. viridis

L. wagneri

Physalaemus gracilis

Completed melanonotus distribution data sheets

Distribution notes on the species groups

Distribution of melanonotus group

Underwater Component in the Call of the Frog Leptodactylus ocellatus, by Alejandro Purgue

Caldwell - Brasilian frog data - NSF reports

J. R. Dixon data on Leptodactylus at El Bagual, Argentina

Chiasmocleis

Call of Eleutherodactylus sp. Brejo da Lapa, WRH 8429

Aborted Leptodactylus pentadactylus biogeography project

Wynn and Heyer - Genetic relatedness within L. fuscus (electronic records)

Bertha Lutz bibliography

Brasil frog literature, notes, etc.

Leptodactylus loan sheets - Completed

Leptodactylus types - Negatives and miscellaneous

Current loans

Leptodactylus labialis - Distribution

Pentadactylus group - Locality information

Rana macrocephala Wied (=Ceratrophrys)

The Society of the Sigma Xi certificate, 1966

Phi Sigma Society certificate, 1964

"A Diatom Is Two Atoms Working Together" - A collection of classic examination answers of biology 100 and 106 students

Leptodactylus - Miscellaneous correspondence, 1963-1968

Completed Pacific Lutheran University loan invoices 1-25

Leptodactylus correspondence and miscellaneous (3 folders)

Leptodactylus data for phylogenetic analysis

Leptodactylus holdings at University of Texas, Arlington, 2006

Anais da Academia Brasileira de Ciencias, 76(2), 2004

Heyer bibliography

Heyer, W. R. 1967. A herpetofaunal study of an ecological transect through the Cordillera de Tilaran, Costa Rica. Copeia, 1967: 259-271.

Savage, J. M., and Heyer, W. R. 1967. Variation and distribution in the tree-frog genus Phyllomedusa in Costa Rica, Central America. Beitrage zur Neotropischen Fauna, 5: 111-131.

Dixon, J. R., and Heyer, W. R. 1968. Anuran succession in a temporary pond in Colima, Mexico. Bulletin of the Southern California Academy of Sciences, 67: 129-137.

Heyer, W. R. 1968. The proper name for the type-species of the genus Leptodactylus. Copeia, 1968: 160-162.

Heyer, W. R. 1969. The adaptive ecology of the species groups of the genus Leptodactylus (Amphibia, Leptodactylidae). Evolution, 23: 421-428.

Heyer, W. R., and Silverstone, P. A. 1969. The larva of the frog Leptodactylus hylaedactylus (Leptodactylidae). Fieldiana Zoology, 51: 141-145.

Heyer, W. R. 1969. Studies on frogs of the genus Leptodactylus (Amphibia, Leptodactylidae). 5. Taxonomic notes on L. latinasus, rhodonotus, romani and wuchereri. Herpetologica, 25: 1-8.

Heyer, W. R. 1969. Studies on the genus Lepodactylus (Amphibia, Leptodactylidae). 3. A redefinition of the genus Leptodactylus and a description of a new genus of leptodactylid frogs. Contributions in Science, 155: 1-14.

Savage, J. M., and Heyer, W. R. 1969. The tree-frogs (Family Hylidae) of Costa Rica: Diagnosis and distribution. Revista de Biologia Tropical, 16(1): 1-127.

Heyer, W. R. 1970. Studies on the frogs of the genus Leptodactylus (Amphibia: Leptodactylidae). 6. Biosystematics of the melanonotus group. Contributions in Science, 191: 1-48.

Heyer, W. R. 1970. Gliding speeds of Ptychozoon lionatum (Reptilia: Gekkonidae) and Chrysopelea ornata (Reptilia: Colubridae). Herpetologica, 26: 317-319.

Heyer, W. R. 1970. Studies on the genus Leptodactylus (Amphibia: Leptodactylidae). 2. Diagnosis and distribution of the Leptodactylus of Costa Rica. Revista de Biologia Tropical, 16(2): 171-205.

Heyer, W. R. 1971. Mating calls of some frogs from Thailand. Fieldiana Zoology, 58: 61-82.

Heyer, W. R. 1971. Descriptions of some tadpoles from Thailand. Fieldiana Zoology, 58: 83-91.

Heyer, W. R., and Peters, J. A. 1971. The frog genus Leptodactylus in Ecuador. Proceedings of the Biological Society of Washington, 84: 163-170.

Heyer, W. R. 1971. The tadpole of Rana pileata. Herpetological Review, 3(5): 95.

Heyer, W. R. 1971. Leptodactylus labialis. Catalogue of American Amphibians and Reptiles, 104: 1-3.

Heyer, W. R. 1972. A new limbless skink (Reptilia: Scincidae) from Thailand with comments on the generic status of the limbless skinks of Southeast Asia. Fieldiana Zoology, 58: 109-129.

Heyer, W. R. 1972. The status of Leptodactylus pumilio Boulenger (Amphibia, Leptodactylidae) and the description of a new species of Leptodactylus from Ecuador. Contributions in Science, 231: 1-8.

Heyer, W. R. 1973. Systematics of the marmoratus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae). Contributions in Science, 251: 1-50.

Heyer, W. R. 1973. Ecological interactions of frog larvae at a seasonal tropical location in Thailand. Journal of Herpetology, 7(4): 337-361.

Heyer, W. R., and Bellin, M. S. 1973. Ecological notes on five sympatric Leptodactylus (Amphibia, Leptodactylidae) from Ecuador. Herpetologica, 29(1): 66-72.

Heyer, W. R., and Berven, K. A. 1973. Species diversities of herpetofaunal samples from similar microhabitats at two tropical sites. Ecology, 54(3): 642-645.

Heyer, W. R. 1974. Niche measurements of frog larvae from a seasonal tropical location in Thailand. Ecology, 55(3): 651-656.

Heyer, W. R. 1974. Relationships of the marmoratus species group (Amphibia, Leptodactylidae) within the subfamily Leptodactylinae. Contributions in Science, 253: 1-45.

Heyer, W. R. 1974. Vanzolinius, a new genus proposed for Leptodactylus discodactylus (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 87(11): 81-90.

Heyer, W. R., and Diment, J. M. 1974. The karyotype of Vanzolinius discodactylus and comments on usefulness of karyotypes in determining relationships in the Leptodactylus-complex (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 87(29): 327-336.

McDiarmid, R. W., Heyer, W. R., and Weigmann, D. L. 1974. Tadpoles, predation and pond size in the tropics. Herpetological Review, 5(3): 76.

Heyer, W. R. 1975. Adenomera lutzi (Amphibia: Leptodactylidae), a new species of frog from Guyana. Proceedings of the Biological Society of Washington, 88(28): 315-318.

Heyer, W. R., McDiarmid, R. W., and Weigmann, D. L. 1975. Tadpoles, predation and pond habitats in the tropics. Biotropica, 7(2): 100-111.

Heyer, W. R. 1975. A preliminary analysis of the intergeneric relationships of the frog family Leptodactylidae. Smithsonian Contributions to Zoology, 199: 1-55.

Pyburn, W. F., and Heyer, W. R. 1975. Identity and call of the frog, Leptodactylus stenodema. Copeia, 3: 585-587.

Heyer, W. R. 1976. Notes on the frog fauna of the Amazon Basin. Acta Amazonica, 6(3): 369-378.

Heyer, W. R. 1976. Studies in larval amphibian habitat partitioning. Smithsonian Contributions to Zoology, 242: 1-27.

Heyer, W. R. 1976. [Review] "Evolution of the Desert Biota." Copeia, 3: 614-615.

Heyer, W. R. 1976. The presumed tadpole of Paratelmatobius lutzi (Amphibia, Leptodactylidae). Papeis Avulsos de Zoologia, 30(10): 133-135.

Heyer, W. R., and Liem, D. S. 1976. Analysis of the intergeneric relationships of the Australian frog family Myobatrachidae. Smithsonian Contributions to Zoology, 233: 1-29.

Heyer, W. R., and Muedeking, M. H. 1976. Notes on tadpoles as prey for naiads and turtles. Journal of Washington Academy of Sciences, 66(4): 235-239.

Muedeking, M. H., and Heyer, W. R. 1976. Descriptions of eggs and reproductive patterns of Leptodactylus pentadactylus (Amphibia: Leptodactylidae). Herpetologica, 32: 137-139.

Straughan, I. R., and Heyer, W. R. 1976. A functional analysis of the mating calls of the Neotropical frog genera of the Leptodactylus complex (Amphibia, Leptodactylidae). Papeis Avulsos de Zoologia, 29(23): 221-245.

Heyer, W. R. 1977. A discriminant function analysis of the frogs of the genus Adenomera (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 89(51): 581-592.

Heyer, W. R. 1977. Taxonomic notes on frogs from the Madeira and Purus Rivers, Brasil. Papeis Avulsos de Zoologia, 31(8): 141-162.

Heyer, W. R. 1977. Foam nest construction in the leptodactylid frogs Leptodactylus pentadactylus and Physalaemus pustulosus (Amphibia, Anura, Leptodactylidae). Journal of Herpetology, 11(2): 225-228.

Heyer, W. R. 1978. Systematics of the fuscus group of the frog genus Leptodactylus (Amphibia, Leptodactylidae). National History Museum of Los Angeles County, Science Bulletin, 8(29): 1-85.

Heyer, W. R. 1978. Variation in members of the Centrolenella eurygnatha complex (Amphibia: Centrolenidae) from Serra do Mar and Serra da Mantiqueira, Brasil. Papeis Avulsos de Zoologia, 32(2): 15-33.

Heyer, W. R. 1978. [Review] "A Biogeographical Analysis of the Chihuahuan Desert through Its Herpetofauna." Herpetological Review, 9(2): 50-51.

Heyer, W. R. 1979. Annual variation in larval amphibian populations within a temperate pond. Journal of the Washington Academy of Sciences, 69(2): 65-74.

Heyer, W. R. 1979. Systematics of the pentadactylus species group of the frog genus Leptodactylus (Amphibia: Leptodactylidae). Smithsonian Contributions to Zoology, 301: 1-43.

Heyer, W. R., and Crombie, R. I. 1979. Natural history notes on Craspedoglossa stejnegeri and Thoropa petropolitana (Amphibia: Salientia, Leptodactylidae). Journal of the Washington Academy of Sciences, 69(1): 17-20.

Heyer, W. R., and Mello, C. C. 1979. Descriptions of the advertising calls of Cycloramphus asper and Cycloramphus dubius (Amphibia: Leptodactylidae). Papeis Avulsos de Zoologia, 32(15): 193-200.

Heyer, W. R. 1980. The calls and taxonomic positions of Hyla giesleri and Ololygon opalina (Amphibia: Anura: Hylidae). Proceedings of the Biological Society of Washington, 93(3): 655-661.

Maxson, L. R., Heyer, W. R., and Maxson, R. D. 1981. Phylogenetic relationships of the Brasilian leptodactylid frog genera Craspedoglossa, Cycloramphus and Zachaenus. Experientia, 37(2): 144-145.

Drewry, G. E., Heyer, W. R., and Rand, A. S. 1982. A functional analysis of the complex call of the frog Physalaemus pustulosus. Copeia, 3: 636-645.

Heyer, W. R. 1982. Two new species of the frog genus Hylodes from Caparao, Minas Gerais, Brasil (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 95(2): 377-385.

Heyer, W. R., Daugherty, C. H., and Maxson, L. R. 1982. Systematic resolution of the genera of the Crinia complex (Amphibia: Anura: Myobatrachidae). Proceedings of the Biological Society of Washington, 95(3): 423-427.

Heyer, W. R., and Maxson, L. R. 1982. Distributions, relationships, and zoogeography of lowland frogs: The Leptodactylus complex in South America, with special reference to Amazonia. In: Prance, G. T. (ed.), Biological diversification in the tropics. New York: Columbia University Press, pp. 375-388.

Heyer, W. R., and Maxson, L. R. 1982. Neotropical frog biogeography: Paradigms and problems. American Zoologist, 22(2): 397-410.

Maxson, L. R., and Heyer, W. R. 1982. Leptodactylid frogs and the Brasilian Shield: an old and continuing adaptive relationship. Biotropica, 14(1): 10-15.

Crombie, R. I., and Heyer, W. R. 1983. Leptodactylus longirostris (Anura: Leptodactylidae): Advertisement call, tadpole, ecological and distributional notes. Revista Brasileira de Biologia, 43(3): 291-296.

Heyer, W. R. 1983. Variation and systematics of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae). Arquivos de Zoologia, 30(4): 235-339.

Heyer, W. R. 1983. Clarification of the names Rana mystacea Spix, 1824, Leptodactylus amazonicus Heyer, 1978 and a description of a new species, Leptodactylus spixi (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 96(2): 270-272.

Heyer, W. R. 1983. Notes on the frog genus Cycloramphus (Amphibia: Leptodactylidae), with descriptions of two new species. Proceedings of the Biological Society of Washington, 96(3): 548-559.

Heyer, W. R., and Maxson, L. R. 1983. Relationships, zoogeography, and speciation mechanisms of frogs of the genus Cycloramphus (Amphibia, Leptodactylidae). Arquivos de Zoologia, 30(5): 341-373.

Heyer, W. R., and Pyburn, W. F. 1983. Leptodactylus riveroi, a new frog species from Amazonia, South America (Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 96(3): 560-566.

Wassersug, R. J., and Heyer, W. R. 1983. Morphological correlates of subaerial existence in leptodactylid tadpoles associated with flowing water. Canadian Journal of Zoology, 61(4): 761-769.

Heyer, W. R. 1984. The systematic status of Adenomera griseigularis Henle, with comments on systematic problems in the genus Adenomera (Amphibia: Leptodactylidae). Amphibia-Reptilia, 5(2): 97-100.

Heyer, W. R. 1984. Variation, systematics, and zoogeography of Eleutherodactylus guentheri and closely related species (Amphibia: Anura: Leptodactylidae). Smithsonian Contributions to Zoology, 402: 1-42.

Heyer, W. R. 1984. [Review] Advances in Herpetology and Evolutionary Biology: Essays in Honor of Ernest E. Williams. Rhodin, Anders G. J., and Miyata, K. Science, 223(4632): 162.

Heyer, W. R. 1985. Taxonomic and natural history notes on frogs of the genus Centrolenella (Amphibia: Centrolenidae) from southeastern Brasil and adjacent Argentina. Papeis Avulsos de Zoologia, 36(1): 1-21.

Heyer, W. R. 1985. New species of frogs from Boraceia, Sao Paulo, Brazil. Proceedings of the Biological Society of Washington, 98(3): 657-671.

Vanzolini, P. E., and Heyer, W. R. 1985. The American herpetofauna and the interchange. Topics in Geobiology, 4: 475-487.

Heyer, W. R., and Cocroft, R. B. 1986. Descriptions of two new species of Hylodes from the Atlantic forests of Brazil (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 99(1): 100-109.

Cocroft, R. B., and Heyer, W. R. 1988. Notes on the frog genus Thoropa (Amphibia: Leptodactylidae) with a description of a new species (Thoropa saxatilis). Proceedings of the Biological Society of Washington, 101(1): 209-220.

Heyer, W. R. 1988. A notable collection of Cycloramphus (Amphibia, Leptodactylidae) from Bahia, Brazil, with a description of a new species (Cycloramphus migueli). Proceedings of the Biological Society of Washington, 101(1): 151-154.

Heyer, W. R. 1988. On frog distribution patterns east of the Andes. In: Vanzolini, P. E., and Heyer, W. R. (eds.), Proceedings of a Workshop on Neotropical Distribution Patterns Held 12-16 January 1987. Rio de Janeiro: Academia Brasileira de Ciencias, pp. 245-273.

Heyer, W. R., Rand, A. S., Cruz, C. A. G., and Peixoto, O. L. 1988. Decimations, extinctions, and colonizations of frog populations in southeast Brazil and their evolutionary implications. Biotropica, 20(3): 230-235.

Heyer, W. R., and Vanzolini, P. E. 1988. Introduction and summary of a workshop on Neotropical distribution patterns held 12-16 January 1987. In: Vanzolini, P. E., and Heyer, W. R. (eds.), Proceedings of a Workshop on Neotropical Distribution Patterns Held 12-16 January 1987. Rio de Janeiro: Academia Brasileira de Ciencias, pp. 7-18.

Maxson, L. R., and Heyer, W. R. 1988. Molecular systematics of the frog genus Leptodactylus (Amphibia: Leptodactylidae). Fieldiana Zoology, 41: 1-13.

Wassersug, R. J., and Heyer, W. R. 1988. A survey of internal oral features of leptodactyloid larvae (Amphibia: Anura). Smithsonian Contributions to Zoology, 457: 1-99.

Heyer, W. R., and Wolf, A. J. 1989. Physalaemus crombiei (Amphibia, Leptodactylidae), a new frog species from Espirito Santo, Brazil with comments on the Physalaemus-signifer group. Proceedings of the Biological Society of Washington, 102(2): 500-506.

Heyer, W. R., Rand, A. S., Cruz, C. A. G., Peixoto, O. L., and Nelson, C. E. 1990. Frogs of Boraceia. Arquivos de Zoologia, 31(4): 231-410.

Heyer, W. R., and Hardy, L. M. 1991. A new species of frog of the Eleutherodactylus lacrimosus assembly from Amazonia, South America (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 104(3): 436-447.

Dubois, A., and Heyer, W. R. 1992. Leptodactylus labialis, the valid name for the American Whitelipped Frog (Amphibia: Leptodactylidae). Copeia, 2: 584-585.

Heyer, W. R., and Perez, L. G. 1993. Description of the advertisement call and resolution of the systematic status of Leptodactylus gracilis delattini Muller, 1968 (Amphibia, Leptodactylidae). Proceedings of the Biological Society of Washington, 106(1): 51-56.

Heyer, W. R., Donnelly, M. A., McDiarmid, R. W., Hayek, Lee-Ann C., and Foster, M. S. 1994. Measuring and Monitoring Biological Diversity: Standard Methods for Amphibians. Smithsonian Institution Press, Washington and London. 364 pages.

Heyer, W. R., Donnelly, M. A., McDiarmid, R. W., Hayek, Lee-Ann C., and Foster, M. S. 1994. Medicion y Monitoreo de la Diversidad Biologica: Metodos estandarizados para Anfibios. Editorial Universitaria de la Patagonia, 349 pages.

Heyer, W. R. 1994. Variation within the Leptodactylus podicipinus-wagneri complex of frogs (Amphibia: Leptodactylidae). Smithsonian Contributions to Zoology, 546: 1-124.

Heyer, W. R. 1994. Hyla benitezi (Amphibia: Anura: Hylidae): First record for Brazil and its biogeographical significance. Journal of Herpetology, 28(4): 497-499.

Cardoso, A. J., and Heyer, W. R. 1995. Advertisement, aggressive, and possible seismic signals of the frog Leptodactylus syphax (Amphibia, Leptodactylidae). Alytes, 13(2): 67-76.

Heyer, W. R. 1995. South American rocky habitat Leptodactylus (Amphibia: Anura: Leptodactylidae) with description of two new species. Proceedings of the Biological Society of Washington, 108(4): 695-716.

Heyer, W. R., and Gascon, C. 1995. Collection notes and call characteristics for Ischnocnema quixensis and Phyzelaphryne miriamae (Amphibia: Anura; Leptodactylidae). Journal of Herpetology, 29(2): 304-307.

Heyer, W. R., and Morales, V. R. 1995. The advertisement call of the leptodactylid frog Leptodactylus griseigularis. Amphibia-Reptilia, 16(1): 91-92.

Heyer, W. R. 1996. Vanishing amphibians: a real problem? World Conservation. 1(96): 10-11.

Heyer, W. R., de Sa, R. O., McCranie, J. R., and Wilson, L. D. 1996. Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae): Natural history notes, advertisement call, and relationships. Herpetological Natural History, 4(2): 169-174.

Heyer, W. R., Garcia Lopez, J. M., and Cardoso, A. J. 1996. Advertisement call variation in the Leptodactylus mystaceus species complex (Amphibia: Leptodactylidae) with a description of a new sibling species. Amphibia-Reptilia, 17(1): 7-31.

Halliday, T. R., and Heyer, W. R. 1997. The case of the vanishing frogs. Technology Review, 100(4): 56-63.

Heyer, W. R. 1997. Geographic variation in the frog genus Vanzolinius (Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 110(3): 338-365.

Savage, J. M., and Heyer, W. R. 1997. Digital webbing formulae for anurans: a refinement. Herpetological Review, 28(3): 131.

Heyer, W. R. 1998. The relationships of Leptodactylus diedrus (Anura, Leptodactylidae). Alytes, 16(1-2): 1-24.

Kress, W. J., Heyer, W. R., Acevedo-Rodriguez, P., Coddington, J. A., Cole, D., Erwin, T. L., Meggers, B. J., Pogue, M. G., Thorington, R. W., Jr., Vari, R. P., Weitzman, Marilyn J., and Weitzman, S. H. 1998. Amazonian biodiversity: Assessing conservation priorities with taxonomic data. Biodiversity and Conservation, 7(12): 1577-1587.

Heyer, W. R., Coddington, Jonathan A., Kress, W. John, Acevedo-Rodriguez, Pedro, Cole, Daniel G., Erwin, Terry L., Meggers, Betty J., Pogue, Michael G., Thorington, R. W., Jr., Vari, R. P., Weitzman, M. J., and Weitzman, S. H. 1999. Amazonian biotic data and conservation decisions. Ciencia e Cultura, 51(5-6): 372-385.

Heyer, W. R., and Munoz, A. M. 1999. Validation of Eleutherodactylus crepitans Bokermann, 1965, notes on the types and type locality of Telatrema heterodactylum Miranda-Ribeiro, 1937, and description of a new species of Eleutherodactylus from Mato Grosso, Brazil (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 112(1): 1-18.

Heyer, W. R. 2000. The lectotype of Cystignathus hylodes Reinhardt and Lutken, 1862. Journal of Herpetology, 34(1): 150-153.

Heyer, W. R., and Morato de Carvalho, C. 2000. Calls and calling behavior of the frog Leptodactylus natalensis (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 113(1): 284-290.

Heyer, W. R., and Morato de Carvalho, C. 2000. The enigmatic advertisement call of Eleutherodactylus ramagii (Amphibia: Anura: Leptodactylidae). Amphibia-Reptilia, 21(1): 117-121.

Heyer, W. R., and Thompson, A. S. 2000. Leptodacylus rugosus Noble. Catalogue of American Amphibians and Reptiles. 708: 1-5.

Heyer, W. R. 2000. Amphibian declines and extinctions: The situation in Latin America. Ecologia Latinoamericana. 47-55.

Carey, C., Heyer, W. R., Wilkinson, J., Alford, R. A., Arntzen, J. W., Halliday, T., Hungerford, L., Lips, K. R., Middleton, E. M., Orchard, S. A., and Rand, A. S. 2001. Amphibian declines and environmental change: Use of remote-sensing data to identify environmental correlates. Conservation Biology, 15(4): 903-913.

Hayek, Lee-Ann C., Heyer, W. R., and Gascon, C. 2001. Frog morphometrics: A cautionary tale. Alytes, 18(3-4): 153-177.

Heyer, W. R., and Heyer, M. M. 2001. Leptodactylus lithonaetes Heyer. Catalogue of American Amphibians and Reptiles, 723: 1-3.

Wynn, Addison H., and Heyer, W. R. 2001. Do geographically widespread species of tropical amphibians exist? An estimate of genetic relatedness within the Neotropical frog Leptodactylus fuscus (Schneider 1799) (Anura Leptodactylidae). Tropical Zoology, 14: 255-285.

Heyer, W. R. 2002. Leptodactylus fragilis, the valid name for the Middle American and northern South American white-lipped frog (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 115(2): 321-322.

Heyer, W. R., and Caramaschi, U. 2002. Comment on the proposed conservation of the specific name of Leptodactylus chaquensis Cei, 1950 (Amphibia, Anura). Bulletin of Zoological Nomenclature, 59(1): 44-45.

Heyer, W. R., de Sa, R. O., and Muller, S. 2002. Leptodactylus silvanimbus McCranie, Wilson, and Porras. Catalogue of American Amphibians and Reptiles, 743: 1-3.

Heyer, W. R., and Heyer, M. M. 2002. Leptodactylus elenae Heyer. Catalogue of American Amphibians and Reptiles, 742: 1-5.

Heyer, M. M., Heyer, W. R., Spear, S., and de Sa, R. O. 2003. Leptodactylus mystacinus (Burmeister) Mustached Frog. Catalogue of American Amphibians and Reptiles. 767: 1-11.

Heyer, W. R., and Junca, Flora A. 2003. Leptodactylus caatingae, a new species of frog from eastern Brazil (Amphibia: Anura: Leptodactylidae). Proceedings of the Biological Society of Washington, 116(2): 317-329.

Heyer, W. R., and Reid, Yana R. 2003. Does advertisement call variation coincide with genetic variation in the genetically diverse frog taxon currently known as Leptodactylus fuscus (Amphibia: Leptodactylidae)? Anais da Academia Brasileira de Ciencias, 75(1): 39-54.

Heyer, W. R. 2004. Paulo Emilio Vanzolini. Copeia, 2004(1): 184-189.

Heyer, W. R., and Heyer, M. M. 2004. Leptodactylus furnarius Sazima and Bokermann cerrado oven frog. Catalogue of American Amphibians and Reptiles, 785: 1-5.

Brandao, R. A., and Heyer, W. R. 2005. The complex calls of Leptodactylus pustulatus (Amphibia, Anura, Leptodactylidae). Amphibia-Reptilia, 26(4): 566-570.

Caramaschi, U., de Sa, R. O., and Heyer, W. R. 2005. Common names for the frog genus Leptodactylus (Amphibia, Anura, Leptodactylidae). Herpetological Review, 36(2): 119-120.

de Sa, R. O., Heyer, W. R., and Camargo, A. 2005. A phylogenetic analysis of Vanzolinius Heyer, 1974 (Amphibia, Anura, Leptodactylidae): Taxonomic and life history implications. Arquivos do Museu Nacional, 63(4): 707-726.

de Sa, R. O., Heyer, W. R., and Camargo, A. 2005. Are Leptodactylus didymus and L. mystaceus phylogenetically sibling species (Amphibia, Anura, Leptodactylidae)? Herpetologia Petropolitana, Russian Journal of Herpetology, 12: 90-92.

Hayek, Lee-Ann C., and Heyer, W. R. 2005. Determining sexual dimorphism in frog measurement data: Integration of statistical significance, measurement error, effect size and biological significance. Anais da Academia Brasileira de Ciencias, 77(1): 45-76.

Heyer, W. R., de Sa, R. O., and Muller, S. 2005. On the enigmatic distribution of the Honduran endemic Leptodactylus silvanimbus (Amphibia: Anura: Leptodactylidae). In: Donnelly, M. A., Crother, B. I., Guyer, C., Wake, M. H., and White, M. E. (eds.), Ecology and Evolution in the Tropics: A Herpetological Perspective. Chicago: University of Chicago Press, pp. 81-101.

Heyer, W. R., and Murphy, J. B. 2005. Declining Amphibian Populations Task Force. In: Lannoo, Michael (ed.), Amphibian Declines: The Conservation Status of United States Species. Berkeley, CA: University of California Press, pp. 17-21.

Heyer, W. R. 2005. Leptodactylus fragilis Brocchi, 1877. White-lipped frog. In: Lannoo, M. (ed.), Amphibian Declines: The Conservation Status of United States Species. Berkeley, CA: University of California Press, pp. 500-501.

Heyer, W. R., and Crombie, R. I. 2005. Leptodactylus lauramiriamae, a distinctive new species of frog (Amphibia: Anura: Leptodactylidae) from Rondonia, Brazil. Proceedings of the Biological Society of Washington, 118(3): 590-595.

Heyer, W. R. 2005. Variation and taxonomic clarification of the large species of the Leptodactylus pentadactylus species group (Amphibia: Leptodactylidae) from Middle America, northern South America, and Amazonia. Arquivos de Zoologia, 37(3): 269-348.

Heyer, W. R., de Sa, R. O., and Rettig, A. 2005. Sibling species, advertisement calls, and reproductive isolation in frogs of the Leptodactylus pentadactylus species cluster (Amphibia, Leptodactylidae). Herpetologia Petropolitana, Russian Journal of Herpetology, 12: 35-39.

Camargo, Arley, de Sa, R. O., and Heyer, W. R. 2006. Phylogenetic analyses of mtDNA sequences reveal three cryptic lineages in the widespread neotropical frog Leptodactylus fuscus (Schneider, 1799) (Anura, Leptodactylidae). Biological Journal of the Linnean Society, 87(2): 325-341.

Heyer, W. R. 2006. Obituary: In celebration of A. Stanley Rand (1932-2005). Phyllomedusa, 5(1): 8-10.

Heyer, W. R., and Heyer, M. M. 2006. Leptodactylus knudseni Heyer, Knudsen's thin-toed frog. Catalogue of American Amphibians and Reptiles, 807: 1-12.

Heyer, W. R., and Heyer, M. M. 2006. Leptodactylus natalensis A. Lutz, bubbling frog. Catalogue of American Amphibians and Reptiles, 808: 1-5.

Heyer, M. M., Heyer, W. R., and de Sa, R. O. 2006. Leptodactylus fragilis (Brocchi) White-lipped Thin-toed Frog. Catalogue of American Amphibians and Reptiles, 830: 1-26.

Heyer, W. R., and Scott, N. J., Jr. 2006. The advertisement call of Leptodactylus laticeps (Amphibia, Anura, Leptodactylidae): predatory aural luring? Herpetological Natural History, 9(2): 189-194.

Heyer, W. R., Caramaschi, U., and de Sa, R. O. 2006. Case 3323. Rana ocellata Linnaeus, 1758 (currently Leptodactylus ocellatus; Amphibia, Anura): proposed conservation of usage of the specific name by the designation of a neotype. Bulletin of Zoological Nomenclature, 63(3): 184-186.

Yanek, K., Heyer, W. R., and de Sa, R. O. 2006. Genetic resolution of the enigmatic Lesser Antillean distribution of the frog Leptodactylus validus (Anura, Leptodactylidae). South American Journal of Herpetology, 1(3): 192-201.

Ponssa, M. L., and Heyer, W. R. 2007. Osteological characterization of four putative species of the genus Adenomera (Anura: Leptodactylidae), with comments on intra- and interspecific variation. Zootaxa, 1403: 37-54.

Bolanos, F., Castro, F., De la Riva, I., Grant, T., Hedges, B., Heyer, W. R., Ibanez D., Roberto, La Marca, E., Lavilla, E., Leite Silvano, D., Lotters, S., Parra Olea, G., Reichle, S., Reynolds, Robert P., Rodriguez, L., Santos Barrera, G., Scott, N., Ubeda, C., Veloso, A., Wilkinson, M., and Young, B. 2008. Amphibians of the Neotropical Realm. In: Stuart, S. N., Hoffmann, M., Chanson, J. S., Cox, N. A., Berridge, R. J., Ramani, P., and Young, B. E. (eds.), Threatened Amphibians of the World. Barcelona: Lynx Edicions, pp. 92-105.

Heyer, W. R., Heyer, M. M., and de Sa, R. O. 2008. Leptodactylus cunicularius Sazima and Bokermann, rabbit-burrow frog. Catalogue of American Amphibians and Reptiles, 845: 1-5.

Heyer, W. R. 2008. Can long-established scientific societies survive the digital age? Journal of the Washington Academy of Sciences, 94(2): 17-32.

Camargo, Arley, Heyer, W. R., and de Sa, R. O. 2009. Phylogeography of the frog Leptodactylus validus (Amphibia: Anura): Patterns and timing of colonization events in the Lesser Antilles. Molecular Phylogenetics and Evolution, 53: 571-579.

Fernandez, M., Cole, D. G., Heyer, W. R., Reichle, Steffen, and de Sa, R. O. 2009. Predicting Leptodactylus (Amphibia, Anura, Leptodactylidae) distributions: Broad-ranging versus patchily distributed species using a presence-only environmental niche modeling technique. South American Journal of Herpetology, 4(2): 103-116.

Heyer, M. H., Heyer, W. R. and de Sa, R. O. 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Amphibia, Anura, Leptodactylidae). Volume 1. References. Smithsonian Herpetological Information Service, 1: 1-677.; Heyer, M. H., Heyer, W. R., and de Sa, R. O. 2009. Bibliography of the frogs of the Leptodactylus clade - Adenomera, Hydrolaetare, Leptodactylus, Lithodytes (Amphibia, Anura, Leptodactylidae). Volume 2. Indices. Smithsonian Herpetological Information Service, 2: 1-167. (electronic records)

Heyer, W. R., and Barrio-Amoros, C. L. 2009. The advertisement calls of two sympatric frogs, Leptodactylus lithonaetes (Amphibia: Anura: Leptodactylidae) and Pristimantis vilarsi (Amphibia: Anura: Strabomantidae). Proceedings of the Biological Society of Washington, 122(3): 282-291.

Heyer, W. R., and Giaretta, A. A. 2009. Advertisement calls, notes on natural history, and distribution of Leptodactylus chaquensis (Amphibia: Anura: Leptodactylidae) in Brasil. Proceedings of the Biological Society of Washington, 122(3): 292-305.

Furness, A. I., McDiarmid, R. W., Heyer, W. R., and Zug, G. R. 2010. Oviduct modifications in foam-nesting frogs, with emphasis on the genus Leptodactylus (Amphibia, Leptodactylidae). South American Journal of Herpetology, 5(1): 13-29.

Heyer, W. R., Heyer, M. M., and de Sa, R. O. 2010. Leptodactylus savagei Heyer Savage's thin-toed frog. Catalogue of American Amphibians and Reptiles, 867: 1-19.

Heyer, W. R., Heyer, M. M., and de Sa, R. O. 2010. Leptodactylus syphax Bokermann Burgundy thin-toed frog. Catalogue of American Amphibians and Reptiles, 868: 1-9.

Lavilla, E. O., Langone, J. A., Caramaschi, U., Heyer, W. R., and de Sa, R. O. 2010. The identification of Rana ocellata Linnaeus, 1758. Nomenclatural impact on the species currently known as Leptodactylus ocellatus (Leptodactylidae) and Osteopilus brunneus (Gosse, 1851) (Hylidae). Zootaxa, 2346: 1-16.

Ponssa, M. L., Heyer, W. R., and de Sa, R. O. 2010. Phylogeny of the genus Leptodactylus (Anura, Leptodactylidae) based on characters of morphology and mode of life. Cladistics, 26(2): 222.

Heyer, W. R., and de Sa, R. O. 2011. Variation, Systematics, and Relationships of the Leptodactylus bolivianus Complex (Amphibia: Anura: Leptodactylidae). Smithsonian Contributions to Zoology, 635: 1-58.

Heyer, M. M., Heyer, W. R., and de Sa, R. O. 2011. Leptodactylus pentadactylus (Laurenti) smoky jungle frog. Catalogue of American Amphibians and Reptiles, 887: 1-48.

Heyer, W. R., and Heyer, M. M. 2012. Systematics, distribution, and bibliography of the frog Leptodactylus validus Garman, 1888 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 125(3): 276-294.

Heyer, W. R. 2013. Paulo Emilio Vanzolini, 25 April 1924-30 April 2013. Herpetological Review, 44(3): 362-364.

Heyer, W. R., and Heyer, M. M. 2013. Systematics, distribution, and bibliography of the frog Leptodactylus insularum Barbour, 1906 (Amphibia: Leptodactylidae). Proceedings of the Biological Society of Washington, 126(3): 204-233.

Heyer, W. R. 2014. Morphological analyses of frogs of the Leptodactylus latrans complex (Amphibia, Leptodactylidae) from selected localities in South America. Proceedings of the Biological Society of Washington, 126(4): 369-378.